The Ancestor of Haplogroup G-Z6748, the Terps, Transport Corridors and Landscape Archaeology – Part Eight

This is the eighth part of long story about a 2,850 year gap or absence of documented YDNA haplogroups in the Griff(is)(es)(ith) genetic YDNA paternal line. Various aspects of this gap have been discussed in the prior seven parts of the story. The gap started with the most common recent ancestor associated with haplogroup G-FGC7516 who was born around 2200 BCE. The next documented genetic ancestor in the Griff(is)(es)(ith) YDNA line is an ancestor associated with the G-Z6748 haplogroup. This gap of undocumented YDNA ancestors represents about 95 generations. It is a relatively big gap that spans a migratory path in a period of wide ranging changes in the environment as well as the social fabric of the landscape in northwestern Europe.

This part of the multi-part story (the eighth part) and the next part (the ninth and final part) focuses on a discussion about the ancestor associated with the G-Z6748 haplogroup and the undocumented generations that may have lived immediately before or after his life.

Specifically this part of the story focuses on the environmental influences and possible soecific migratory paths that might be associated with the generations on the tail end of this phylogenetic gap. The final, ninth part of the story focuses on the possible indigenous socio-cultural groups that might have been associated with these YDNA generations.

As discussed in part six of this story, the estimated possible migratory paths of the approximately 95 undocumented generations that are associated with this phylogenetic gap significantly widens as each successive generation approached and passed through the area that is presently known as the Rhine Meuse delta region. The eventual endpoint of this migratory path is where and when the ancestor associated with haplogroup G-Z6748 may have lived.

The possible migratory paths of the Griff(is)(es)(ith) YDNA genetic line were Influenced in varying degrees by the geographical and environmental influences impacting changes in the Rhine, Meuse, and other river watershed areas. As generations of the Griff(is)(es)(ith) YDNA lineage migrated northwestward through the lower Rhine and delta region, the marine activity on the north and west coastline and the changing terrain of the peat lowlands, increasingly played an influencing role on their migratory route.

In the late Roman and medieval periods, large parts of the western and northern Netherlands were covered by extensive peat bogs that changed over time. Research increasingly shows that major geographical changes (e.g., changing river courses, an increase of flooded areas and wet areas) occurred during the transition from the late-Roman (around 270 to 450 CE) to early-medieval periods (roughly 450 to 1050 CE). [1] Coinciding with these landscape changes, archaeological evidence in the modrn day Netherlands area points to a severe demographic decline as well as changes in settlement patterns and land use during this period. [2]

The first millennium AD encompasses the Roman period (12 BC to AD 450) and the Early Middle Ages (AD 450 to 1050). In the Netherlands, this millennium saw population growth, steep decline and subsequent revival. In addition, many changes occurred in the physical landscape, marking a transition from a mainly natural prehistorical lowland landscape to an increasingly human-affected landscape.

From the late 3rd century AD, . . . depopulation occurred, coinciding with the collapse of the Western Roman Empire and the large-scale migration of tribes throughout NW Europe. This period has traditionally been referred to as the ‘Dark Ages’ . This term on the one hand refers to a period of cultural decline and disorder, and on the other hand it is used for periods in general from which little information is available.[3]

The Historical and Geographical Context of the Most Recent Common Ancestor (MRCA) of Haplogroup G-Z6748

As depicted in illustration one, it is estimated that the ancestor that marks the endpoint of this phylogenetic gap, a male descendant who represents the G-Z6748 haplogroup, was born around 668 CE. There is a 95 percent chance of certainty that he was born within a roughly six hundred year time span, 380 CE to 908 CE. There is a 68 percent chance that he was born between 524 CE and 792 CE, roughly a 275 year range (see illustration two). [4]

Illustration One: Scientific Details of the G-Z6748 Haplogroup

Click for Larger View | Source: FamilyTreeDNA,Scientific Details on Haplogroup G-Z6748, https://discover.familytreedna.com/y-dna/G-Z6748/scientific

Harm Jan Pierik, a geographer who studies the geology and landscape evolution of the lowland areas, provides an informative illustration that situates two paleogeographical maps in a timeline based on five different time axes: general archaeological time periods, socio-cultural time periods, reforestation and deforestation cycles, climatic periods and geomorphological changes between 100 BCE and 1200 CE (see illustration two).

Illustration Two: Palaeogeographical maps of 100 CE and 800 CE of Netherlands

Click for larger View | Source: Fig. 2. Palaeogeographical maps of (A) AD 100 and (B) AD 800 in Pierik HJ. Landscape changes and human – landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

This illustration provides a wealth of summary and graphic information on the Netherlands in the first millennium CE. The Dutch landscape shifted from mostly natural to an heavily human-altered environment. Changes, however, varied by region due to the nature of the local geography and people’s actions like farming, draining land, and cutting peat for building supplies and fire. Humans slowly weakened landscapes through everyday land use, until big events like storms tipped them into lasting new shapes—worst in peat coasts, milder in rivers and sands areas. By 1000 CE, people had unintentionally remade the Netherlands’ lowlands.

If we assume that the most recent common ancestor of haplogroup G-Z-6748 was born roughly between 525 and 800 CE (within the 68 percent statistical confidence interval reflected in illustration one), we can get an inkling of the general historical context and the physical circumstances he as well as immediate preceding generations experienced when they lived. Illustration three highlights when the Most Recent Common Ancestor (MRCA) of G-Z6748 and immediate generations lived in context of the five time lines found in Harm Jan Pierik’s illustration.

Illustration Three: Most Recent Common Ancestor of G-Z6748 and Five Time Periods

Click for larger View | Source: Part of Fig. 2. Palaeogeographical maps of (A) AD 100 and (B) AD 800 in Pierik HJ. Landscape changes and human – landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

The ancestor lived in what is known as ‘period B’ of the early medieval archaeoogical period. This early medieval period (roughly the fifth through tenth centuries) in Europe, is called the “Dark Ages” and saw the fall of Rome, migrations of various indigenous groups (referred to as ‘the Migration Period [5] ), and the formation of new kingdoms like Frankish Merovingians in the mid to south region, the Frisians in the north and Anglo-Saxons in England. [6] It was a period characterized by fragmented power, cultural shifts, and the rise of Christianity, with later centuries showing increasing stability and development before the High Middle Ages. [7]

Another rendition of the five time lines in context of when the MRCA of G-Z6748 lived is provided in illustration four below. Illustration four is from a study by Rowin J. van Lanen who presents the combined results of several mutlidisciplinary studies, including Pierik’s, that developed landscape-archaeological models of this time period. These models spatially analyze natural and cultural dynamics in five manifestations: route networks, long-distance transport corridors, settlement patterns, palaeodemographics and land-use systems. Van Lenen’s summary study basically ‘repackages’ Pierik’s illusration into a slightly different graphic portrayal. [8]

Illustration Four: The MRCA of G-Z6748 in Context of Cultural and Natural Dynamics During the First Millennium AD

Click for Larger View | Source: Modified version of Figure 2 in R.J. van Lanen. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences, Volume 99, e14. https://doi.org/10.1017/njg.2020.12

Similar to illustration three, van Lanen’s depiction of the time period suggests that the MRCA of G-Z6748 lived during a period when there was low population density during the Merovingian era that witnessed a period of reforestation. He also lived in the Dark Ages Cold Period (DACP) was a time of widespread cold, particularly in the Northern Hemisphere, roughly from AD 400 to 765 CE, following the Roman Warm Period and overlapping with the Late Antique Little Ice Age (LALIA) around AD 536-660 AD. This cold period was likely triggered by massive volcanic eruptions and low solar activity. [9]

During this period of time, generations of ancesters preceding the MRCA of haplogroup G-Z6748 may have lived in the central area of the lowlands. It is also possible that the MRCA of G-Z6748 migrated to the northern area where remnants of his YDNA have been reportedly located. Depending on the rate and course of migration, preceding generations may have lived in the coastal areas of the central delta and river watershed areas of the Meuse and Rhine Rivers.

“Life Experiences” of MRCA of G-Z6748 and Ancestors Living Between 525 and 800 CE

As mentioned there is a 68 percent chance that the MRCA of G-Z6748 was born between 524 CE and 792 CE. This time span of roughly a 275 years represents about 8 generations of YDNA ancesors, one of which represents the MRCA of G-6748. Using this time span, we can recreate a general protrayal of what these generations experienced based on the time line conditions referenced in illustrations three and four and depending on the geographical location of where they might have lived along the migratory path. 

An individual living in the northern coastal Netherlands area between 525 and 800 CE

An individual in the northern Netherlands between 525 and 800 CE would have lived in small, kin-based rural communities in a wet, tidally influenced landscape, with a mixed farming economy and gradually increasing integration into wider Frisian and Frankish trade networks and political structures of emerging elites. Social life was structured around extended households, local assemblies and daily existence was closely shaped by flooding, salt-marsh grazing, and modest climatic variability in the late Holocene North Sea environment. [10]

Much of the northern coastal zone (Friesland, Groningen, north Noord-Holland) consisted of low salt marshes, tidal creeks, and peat hinterlands, with settlements concentrated on artificial dwelling mounds (called terpen/wierden) rising above flood level.​ Periodic storm surges and high tides inundated surrounding fields and pastures, depositing fertile silt but also posing recurrent risks to people and livestock.​ The climate in the North Sea region showed phases of increased wetness without the sustained warmth of the later Medieval Climate Anomaly. Inhabitants experienced a cool-temperate, often damp and windy regime with notable year-to-year variability. [11]

The core economic experience was mixed farming, with a strong emphasis on cattle and sheep grazing the salt marshes, supplemented by arable plots on slightly higher or reclaimed ground.​ Daily labor included tending livestock, managing manure and fodder on restricted dry surfaces, maintaining paths and small embankments, and exploiting marine resources such as fish and perhaps shellfish from creeks and tidal flats.​ From the later sixth to seventh centuries, coastal communities increasingly tapped into regional exchange networks. Some Frisians gained reputations as merchants and mariners moving goods along the North Sea, though most rural inhabitants remained primarily local farmers with occasional surplus from their labor entering trade. [12]

Politically, the region was characterized by early medieval Frisia: a patchwork of local communities and elites along the North Sea coast, described in later written sources as forming a Frisian “kingdom,” but likely experienced locally as a network of kin groups, chiefs, and regional leaders rather than a centralized state.​ Social life was organized around extended households and free farming families, with local assemblies and customary law; later codified as Lex Frisionum under Frankish rule, this legal culture emphasized fines, compensation, and gradations of social status.​ From the seventh to eighth centuries, Frankish expansion brought military pressure and eventual incorporation of much of Frisia into the Carolingian realm, so inhabitants would increasingly encounter Frankish officials, tribute demands, and shifting allegiances of local elites, even as everyday village life remained relatively continuous. [13]

In the earlier part of this period, religious life centered on local pagan cults and rituals, with sanctuaries and offerings embedded in the landscape. Christian missionaries began to work in Frisian territory from the late seventh century onward.​ Over the eighth century, conversion progressed unevenly. Some communities saw churches established on or beside terpen, while others likely maintained older practices for longer, producing a mixed religious experience with new rites layered onto existing customs. [14]

“Frisian” identity in these centuries was situational and relational, emerging in contacts with Franks, Saxons, and North Sea partners. For most individuals, identity would have been anchored first in kin, settlement, and local region, with broader ethnic labels activated in specific legal, military, or trading contexts. [15]

An individual living in the central delta and river watershed area of the Meuse and Rhine Rivers Between 525 and 800 CE

The daily existence of an individual living in the central delta and river watershed area of the Rhine and Meuse rivers was shaped by mixed farming in a wet, flood‑prone delta landscape. They may have witnessed periodic political and military disruptions over control of the Rhine–Meuse system and a slow Christianization that overlaid older regional cults. The central area (roughly the Rhine–Meuse–Waal–IJssel river district around Utrecht, Nijmegen, and the lower Rhine branches) was a low delta with natural levees, crevasse splays, peat bogs, and backswamps. People concentrated on the slightly higher, drier alluvial ridges along the main channels. [16]

Flooding was a recurring fact of life: around the fifth to sixth centuries. The Waal river branch became more dominant, bringing higher and more frequent floods in parts of the delta, though protective natural levees made the landscape relatively resilient compared to more exposed coastal zones.​ For inhabitants, this meant managing arable strips and meadows on limited high ground, coping with occasional inundations and sedimentation, and negotiating access to extensive wetland resources (peat, reeds, fishing, fowling) in surrounding lowlands. [17]

Settlement focused on small farm clusters and villages strung along levees and sand ridges, often reusing or near former Roman sites. Early medieval farms at places like De Geer or near Roomburg show continuity and adaptation of earlier settlement structures in the river landscape. [18] Households practiced mixed farming (cattle, sheep, some pigs and horses, with cereals and other crops on better-drained plots), with daily work dominated by tending stock, maintaining fields and drainage, cutting peat or sods, and using the rivers as movement corridors.

Compared to the terp coast, inhabitants here had more direct contact with inland and southern regions through river traffic. Even ordinary villagers would periodically encounter non‑local goods, styles, and people via markets and itinerant traders. [19]

Politically, this was a contested geographical zone: during parts of the seventh century, Frisian rulers extended power into the central river area, while Merovingian and later Carolingian Frankish kings held key strongholds and centers farther south and east.​Dorestad, near modern Wijk bij Duurstede, grew into a major trading emporium along the Rhine–Lek fork, so people in its hinterland experienced increased demand for surplus produce, craft goods, and transport services, as well as exposure to coinage and long‑distance merchants.​The Frisian–Frankish wars in the seventh to eighth centuries, focused on control of the Rhine delta and brought episodes of campaigning, shifting overlordship, and, for some communities, tribute or military obligations, even if actual battles occurred only intermittently in any given locality. [20]

The central river region retained deep cultural memories of earlier Batavian and Roman cult sites. Early medieval sources and archaeology indicate regional cult places and sacred landscapes persisting into the first Christian centuries.​ Christianization advanced here earlier and more densely than in the northern coastal zone. Utrecht, built on a former Roman fort, became a missionary center, and churches and cemeteries appeared along the river routes from the seventh century onward. For individuals, this likely meant living through a gradual shift from cremation or traditional burial rites to inhumation in Christian graveyards, new ritual calendars, and the growing authority of priests and ecclesiastical institutions layered onto existing kin and local structures. [21]

The Rhine–Meuse corridor linked the North Sea and Zuiderzee routes with central and southern Frankish regions, so central Netherlands inhabitants were embedded in larger regional exchange systems that carried wine, pottery, textiles, slaves, and other goods.​ Over the seventh to eighth centuries, Frankish consolidation and the rise of emporia-like Dorestad drew the area more firmly into the Carolingian world, creating opportunities in trade and craft but also greater exposure to taxation, tolls, and elite power struggles. [22]

The enviromental impacts in the coastal, river delta, and sandy uplands areas

From 500 to 900 CE the northern Netherlands shifted from a largely open peat–marsh and barrier-island coast with dispersed terp settlements to a more fragmented, wetter and increasingly human‑engineered tidal landscape with expanding salt marsh ridges, drowned peat interiors, and more structured terp rows and ‘proto‑dike’ systems. The key processes were gradual: relative sea‑level rise, peat subsidence and erosion, salt‑marsh progradation along the Wadden Sea, and locally intensive reclamation and drainage that both created and destroyed land. [23]

In the coastal lowlands, peat bogs were drained for farming and fuel, causing them to sink and become vulnerable to sea floods from storms. Once saltwater rushed in, it carved channels and piled up mud, turning huge bog areas into salty tidal flats that stayed that way for centuries, making much of the land unlivable. [24]

In the river delta areas of the Meuse and Rhine rivers, deforestation upstream dumped more dirt into rivers like the Rhine and Meuse, while sinking peat downstream helped new river branches form and steal water from old ones. People kept living on higher riverbanks despite more floods, adapting by moving uphill, but the area stayed mostly farmable without total collapse. [25]

In the sandy uplands of the northern coast, sands stayed stable overall, with only small patches blowing around near villages due to tree-cutting and overfarming, especially after 900 CE as population grew. Unlike wetter areas, these ‘hills’ bounced back easily from human changes in land management. [26]

The Migratory Path to the Most Recent Common Ancestor of Haplogroup G-Z6748

This mutli-part story relies on the estimated migratory path of the Griff(is)(es)(ith) genetic paternal ancestors. This estimated path was created through the use of the FamilyTreeDNA’s online GlobeTrekker program. The ‘on-the-fly’ map generation program is an innovative, interactive phylogeographic feature within the Discover™ platform that visualizes ancestral Y-DNA migration routes. 

GlobeTrekker identifies each designated ancestral haplogroup’s probable position and then connects them backward in time, using ‘cost-efficient‘ routes. These ‘most likely’ or ‘cost-efficient’ routes are estimated within corridor paths or bands, capturing the uncertainty inherent in reconstructing prehistorical movement based on environmental and genetic data. The visible ‘corridor bands’ in GlobeTrekker are explicitly tied to ‘likelihood percentages‘. [27]

If we reduce the visual presence of corridor bands as well as the contrasting colors in the GlobeTrekker interactive map and leave only the estimated migratory path, you obtain a map reflected in illustration five below. The migratory line is the reflection of a ‘minimum-cost path’ within those migratory confidence bands.

Illustration Five: Estimated Migratory Path to Ancestor Associated with Haplogroup G-Z6748

Click for Larger View | Source: Migration Route to ancestral haplogroup G-Y132505 from G-Z67487, GlobeTrekker, FamilyTreeDNA, accessed 4 Dec 2025

What is noteworthy, when looking at the above map, is the approximate location of where the ancestor of haplogroup G-Z6748 may have lived based on the estimates derived from the Globetrekker program.. The outline of the north coast in illustration four represents the current contours of the northern coast of the Netherlands. The island just north of the Netherlands coastline is Texel Island. It is an island that is part of the West Frisian Islands, a chain of barrier islands in the North Sea. Texel is the largest of the Frisian Islands, including, Vlieland, Terschelling, Ameland, and Schiermonnikoog. [28]

If we compare the GlobeTrekker map in illustration five with palaeogeographical maps between 50 CE and 1460 CE (see maps A, B, and C, in illustration six below), it is evident that the coastal and deltaic plains during the time of this migration witnessed profound ecological changes. The modern day Texal area was attached to the mainland and was charactacterized by having high dunes surrounded by low dunes, beach ridges and valleys in 50 CE and 750 CE. By 1450 CE, the land has been separated and is an island.

Illustration Six: The Changing Landscape of the Netherlands Between 50 CE and 1450 CE in Comparison with the Migratory Paith of the Griff(is)(es)(ith) YDNA LIne

Click for Larger View | Source: The three maps are modified versions of Fig. 4. Palaeogeographical maps of the Holocene development of the Dutch coastal and deltaic plain in T. de Haasa,b, H.J. Pierika, A.J.F. van der Spek, K.M. Cohen, B. van Maanen, M.G. Kleinhans, Holocene evolution of tidal systems in The Netherlands: Effects of rivers, coastal boundary conditions, eco-engineering species, inherited relief and human interference Earth-Science Reviews 177 (2018) 139–163.

Texel became a distinct island from the North Holland mainland after this ancestor’s existence in 1170 CE due to the devastating All Saints’ Flood. The flood was the result of a massive storm surge that inundated the land, separating it and creating the island’s current form. Before this event, Texel was connected to the mainland. The floodwaters carved channels that isolated it permanently (see illustration seven). [29]

Illustration Seven: Texal Island and the West Frisian Islands

Click for Larger View | Source: Figure 1 in P.C. Vos & E. Knol, Holocene landscape reconstruction of the Wadden Sea area between Marsdiep and Weser,Netherlands Journal of Geosciences — Geologie en Mijnbouw| 94 – 2, 157–183, 2015, https://njgjournal.nl/index.php/njg/article/view/11860/18365

Based on the estimated ‘cost-efficient’ migratory route generated by GlobeTrekker calculations, it is possible that the ancestors of haplogrup G-Z6748 may have migrated to the northwestern point of what we call modern day Netherlands utilizing water and land routes that were used in the Roman and early medieval eras. Illustration eight depicts the general migratory route in context of the paleogeographical characteristics of the environment around 800 CE. It is possible the ancestors of G-Z6748 may have migrated north-westward via water routes, and to a lessor extent roadways, through areas that witnessed environmental changes, such as increased deforestation and geomorphological change in the northern coastal areas. [30]

Illustration Eight: Palaeogeographical Map 800 CE

Cick for Larger View | Source: Modification of Fig. 1 Main landscape changes in the first millennium AD in T. de Haasa,b, H.J. Pierika, A.J.F. van der Spek, K.M. Cohen, B. van Maanen, M.G. Kleinhans, Holocene evolution of tidal systems in The Netherlands: Effects of rivers, coastal boundary conditions, eco-engineering species, inherited relief and human interference Earth-Science Reviews 177 (2018) 139–163.

In the Rhine-Meuse delta, major geomorphological changes [31] occurred during the late Roman Era and the Early Medieval Period. Generations prior to when the most recent common ancestor of G-Z6748 may have migrated along and through the Meuse Rhine watershed area during this time period of geographical change

The Rhine-Meuse delta area was characterized by a relatively high flooding frequency. For example, research conducted by Esther Jansma reconstructs a dense, year‑by‑year history of floods and related hydrological crises in the northwest European Lowlands (mainly the Netherlands) in the first milenium CE. Her research shows that large floods were clustered in particular centuries rather than evenly distributed. These events coincided with major shifts in settlement patterns and river dynamics. [32]

Jansma’s study identifies about 160 to 170 hydrological “events” (floods, prolonged wet episodes) between in the first millenium, of which roughly 20 to 25 qualify as ‘major flooding events’ defined as greater than or equal to 50‑year recurrence‑interval events.​ One third of all major events fell between roughly 185 to 282 CE. Another pronounced cluster occurs in the later sixth and early seventh century, with the event in 602 CE emerging as the single most severe flood of the millennium in the Dutch dataset. The migratory path of YDNA ancestors of G-Z6748 may have been impacted by both of these two clusters of flooding. [33]

Texal, the Terps and the North Coast – The Possible Home of Generations Associated with Haplogroup G-Z6748

From the later third to fourth century through about 800 CE, the northern Netherlands (roughly Friesland, Groningen, northern Drenthe, north Noord-Holland) was occupied first by remnant and returning Frisian/Chauci-related groups. By the seventh to eighth centuries the area is described in Frankish sources as Frisian territory or a Frisian “kingdom.” (More on ‘the Frisian identity’ is discussed in the ninth and final part of this story. ) [34]

The coastal area of the northern Netherlands where the most recent common ancestor of haplogroup G-Z6748 and related generations may have lived is known as the terp region. It was a region where early inhabitants built artificial mounds called terps (or wierden) on natural salt marshes to survive frequent floods from the rising sea and storm surges, creating a unique landscape of mounds scattered across flat, fertile clay plains. (see illustration nine). This ancient flood protection system allowed permanent settlement from around 600 BCE onwards in an otherwise inhospitable tidal zone. [35]

Illustration Nine: Artistic reconstruction of an early medieval terp settlement 

Click for Larger View | Source: Terps: Ancient Dwelling Mounds of the North Sea Coast, Ealdlar, https://ealdlar.com/history/terps

Initially, a terp might be just large enough for a single farmhouse (house terp), but over time these mounds could be expanded and merged. As population grew, families would enlarge the terp or cluster multiple mounds together, eventually forming a larger village terp hosting several households.[36]

Illustration ten provides a paleogeographical map of the Texal Island area around 800 CE. This time period was probably after the time when the ancestor of G-Z6748 lived. If we compare this map with map C in illustration five or the map in illustration two, the area where this ancestor may have lived may have been separated by a channel or series of channels from the mainland.

Illustration Ten: The Texal Area around 800 CE

Click for Larger View | Source: Figure 10 in P.C. Vos & E. Knol, Holocene landscape reconstruction of the Wadden Sea area between Marsdiep and Weser, Netherlands Journal of Geosciences — Geologie en Mijnbouw| 94 – 2, 157–183, 2015, https://njgjournal.nl/index.php/njg/article/view/11860/18365

In the post-Roman and early medieval period, the terp zone was a low, tidally influenced salt-marsh landscape punctuated by densely occupied artificial mounds that concentrated settlement, livestock and infrastructure above the flood level while exploiting the surrounding grazing marshes and creeks. Those mounds grew into complex village platforms within a dynamic coastal system of progradation [37], erosion and episodic abandonment. [38]

The broader terp area consisted of wide, regularly inundated salt-marshes intersected by tidal creeks and channels, with only slight natural levees or sandy ridges offering higher ground. Sea-level fluctuations and storm surges repeatedly reworked these marshes, creating phases of marsh formation, local drowning of the land and sediment build-up that conditioned where terps could be founded and expanded (see illustration eleven). [39]

Illiustration Eleven: Schematic Representation of the Development of a Terp

Click for Larger View | Source: Fig. 1.2. Schematic representation of the development of a terp, in D. Postma , Salt Marsh Architecture, Catalogue, technology and typological development of early medieval turf buildings in the northern coastal area of the Netherlands, Masters Thesis University of Groningen, 2010, https://www.waddenacademie.nl/fileadmin/inhoud/pdf/06-wadweten/Scripties/D._Postma_-_scriptieverkl.pdf

Description of Illustration Ten: 1. First occupation phase on levee (or marsh bar). 2. Formation of house terpen through the accumulation of refuse and intentional raising. 3. Agglomeration of (nuclear) terpen into a larger village terp. 4. Expansion of the terp comes to an end as the salt marsh is dyked in.

From the Iron Age through the early Middle Ages, farms and later villages were raised on fully artificial mounds (terpen / wierden / warften), constructed from clay, sods and refuse to sit just above typical high-tide and surge levels.​ By the early medieval period individual house-terps (10 to 20 meters across) and larger village-terps (several hectares) formed archipelagos of habitation “islands” within the marsh, often enlarged in phases as population and flood risk increased.

Terp habitation in the regions of the western Netherlands occurred only on a small scale; early terps in this area did not develop into the large dwelling mounds that we know from the northern coastal area. Despite obvious similarities in the Holocene development of the southern North-Sea coastal areas, there are considerable differences between underlying geological characteristics of the western Netherlands on the one hand, and the northern Netherlands and northwestern Germany on the other hand. A large part of the western Netherlands is sheltered by a coastal barrier dune system, whereas the more northerly coastal areas of the Netherlands and Germany were an open salt marsh landscape, an intertidal area of the Wadden Sea, prior to the large-scale medieval dike building. ” [40] (See illustration twelve.)

Illustration Twelve: Discovered Archaeological Sites of Terps in the Wadden Sea Area Based on Archaeological Time Period

Click for Larger View | Source: Figure 1 in A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
https://www.sciencedirect.com/science/article/pii/S0964569118307348

The surrounding marshes provided highly productive grazing for cattle, sheep and horses, so terp communities practiced a mixed agro‑pastoral lifestyle with livestock on the low ground and arable plots and buildings concentrated on the mound tops and shoulders (see illustration thirteen). Excavations show the integration of the terp dwellings into wider economic networks. Roman and later imported goods, as well as isotope evidence of marine-influenced diets, indicate exchange and use of both marine and terrestrial resources. [41]

Illustration Thirteen: Digital 3D Reconstruction of the Mound Landscape

Click for Larger View | Source: Ulco Glimmerveen, 3D renders, Art & Illustration, https://gallery.ulco-art.nl/?page_id=162 . See also, Boeles en Van Giffen Speech or profile paper about mounds, Education,https://terpenonderzoek.nl/education/?lang=en

Between the third and fifth centuries many western terps were thinned out or abandoned, while some eastern terp settlements persisted, reflecting combined environmental stress and shifting socio‑political networks across the North Sea zone (see illustration eleven). Early medieval reoccupation and enlargement of terps, together with new mound building further east and south, created a renewed settlement system that reused earlier structures while adapting layout (larger communal platforms, churches, consolidated farmsteads) to changing social organization. [42]

Terpen typically had linear, lane‑based farm arrangements on a raised platform, with three‑aisled byre‑houses (longhouses) laid out in rows or clusters, often evolving over time from dispersed farmsteads to more nucleated village plans (see ilustration twelve). House plans themselves were usually elongated, tripartite buildings where humans and livestock shared one structure in zoned compartments along a central aisle. [43]

Early terpen often began as one or a few farms on a small, roughly oval or irregular mound, with buildings aligned along the mound’s long axis and oriented to exploit prevailing winds and access to creeks.​ As terpen were enlarged, houses, outbuildings and paths formed more structured layouts: parallel building rows, narrow lanes and yard spaces on the crown and shoulders of the mound, sometimes creating a ring‑like or fan‑shaped pattern around a central open area (see illustration ten). [44]

Between 400 and 800 CE, terp settlements shifted from a phase of contraction and partial abandonment to renewed occupation, enlargement and into more complex village terpen, part of an increasingly more complex Frisian coastal social organizational structure (see table one). Over these four centuries, the settlement pattern, internal layout, social role and regional extent of terpen all changed in step with demographic recovery, environmental opportunities and emerging power structures in the region. [45]

Table One: Terp settlements in the Northern Frisian Area Between 400 – 800 CE

Time PeriodDescription of Terp Settlements
4th–5th centuries: contraction and gapsCeramic sequences and mound stratigraphy [46] indicate that many Frisian terpen in the northern coastal area were abandoned or only sparsely occupied by the 4th century, creating a notable habitation gap after the later Roman period.

In some areas new groups later reoccupied deserted mounds, suggesting discontinuity in local communities even where the same physical terpen were reused.
[47]

Frisian longhouses were three‑aisled buildings, usually 12 to 20 meters long and about 5 to 6 meters wide in the early medieval period, divided lengthwise by two internal post‑rows that carried the roof.​ The internal organization of the house consisted of a residential end (living and sleeping area), a central working or entrance zone with hearth(s), and a byre section for cattle and other stock at the opposite end, all aligned in a single linear plan. [48]
6th–7th centuries: reoccupation and growthFrom around 400–600, occupation continues on selected terpen and new small house‑mounds appear, marking a gradual repopulation of the salt‑marsh zone rather than an immediate return to dense pre‑Roman patterns.

As population and herds increased, individual house terpen were enlarged or physically merged, beginning the process by which scattered farms coalesced into larger village terpen. [49]
7th–8th centuries: nucleated village terpenBy the later 7th and 8th centuries, parts of Frisia show a dense network of occupied terpen—up to roughly 1500 mounds—forming a highly structured coastal settlement system tied to maritime and riverine routes.

Internal layouts become more nucleated: longhouses, ancillary buildings and lanes cluster on shared platforms, and some terpen acquire central functions (elite residences, craft zones, later churches) rather than being purely agrarian farm‑islands. [50]

Terps functioned as ‘nodal points’ linked by seasonal trackways on the marsh, small landing places on creeks and, in some areas, raised routes that later underpinned early medieval road systems. [51] The pattern is best characterised as a multi-centered coastal landscape: multiple terp clusters aligned along former creek systems and lagoon rims, tying maritime access, inland peatlands and higher Pleistocene sands into an highly adapted landscape. [52]

Excavations on Texel (e.g. Den Burg [53]) show early medieval occupation with long continuity of terp‑based and salt‑marsh settlement traditions, but the coastal band between Kennemerland and Westfriesland, including Texel, shows archaeological indications of demographic decline around the fourth century and renewed, denser occupation from the sixth to seventh centuries. For the broader northern Netherlands coast, recent ceramic and settlement analyses indicate relatively thin, scattered habitation in the fifth century, followed by clearer Merovingian‑period rural settlement networks embedded in marsh and tidal landscapes from roughly 600 CE onwards. [54]

The Nature of Road and Waterway Travel Routes that Influenced Migratory Conditions Between 400 and 800 CE

Between 400 and 800 CE the Meuse–Rhine watershed and the coastal Netherlands areas saw a shift from a largely routeless, wetland landscape with localized tracks to a strongly river‑dominated long‑distance network in which fluvial and maritime routes carried the bulk of interregional traffic. Land routes relied on persistent sandy‐ridge paths and levee routes rather than on anything like the earlier Roman road system. [55]

Water-related routes must have been limited to past rivers, inland seas, and shorelines. Land routes show more divers (sic) patterns for these two periods, with large parts of the southern and eastern Netherlands appearing to have been either highly or reasonably accessible. Here sandy ridges constituted corridors through marshy parts of the landscape. [56]

The lowlands of the Rhine–Meuse delta and the northern Netherlands remained highly dynamic, with channel shifts, peat expansion and erosion shaping where routes could exist at all.​ Travel networks in this period depended on a mosaic of higher sandy coversands, levees and older beach ridges which provided dry corridors in an otherwise marshy or peat‑dominated terrain. [57]

” (P)eople living here were strongly dependent on local rivers and streams for transport (water-related routes). An interesting exception are the coastal dunes along the North Sea. Both in A.D. 100 and 800, these dunes provided an accessible north–south land connection. The importance of this connection in the Roman period is underpinned by the occurrence of Roman coastal defenses in these areas.[58]

Formal, stone‑paved Roman roads in the Dutch sector of the limes fell out of regular use after the third to fourth centuries. There is no evidence for a comparable engineered road network in the fifth to seventh centuries.​ Studies of “landscape prerequisites” and route persistence over time suggests that early‑medieval long‑distance paths, where present, hugged sandy ridges and river levees, forming ‘loose corridors’ or ‘route zones’ rather than ‘single fixed roads’. These lines of movement later underlie parts of the documented medieval route skeleton.​ [59]

Roads can be defined as narrow, fixed communication and transport lines connecting different places, whereas routes have been characterized as broad and vaguely delimited zones of communication and transport. Almost all Roman and early-medieval routes were unpaved and hence not rigidly anchored in space. Route zones are spatial zones in which, often unpaved, bundles of tracks, paths or roads are located. These zones formed as a result of travellers frequently shifting to adjacent lanes because of e.g. weather conditions or general wear of the carved-in tracks. Although the general orientation of past roads and routes were similar, route networks spatially were more dynamic and therefore they should be regarded as corridors rather than as single lines.[60]

As indicated in part seven of this story, the Roman state and military investments led to a dramatic rise in road and waterway connectivity. This is aparent in the middle to upper Rhine river areas. The construction of Roman roads, waterways, and quays greatly increased mobility and integration, particularly from the middle of the second century onward. However, as one traveled northward toward the lower Rhine and Delta areas and northward, it is apparent that there were fewer roads connecting cities, towns and forts with the rural areas (see illustration nineteen in part seven of this story).

During the RP (Roman Period) an extensive route network developed, connecting many parts of present-day Europe. However in the Netherlands stone-paved Roman roads such as those present in southern Europe are non-existent including the Oude Rijn (The Old Rhine) . . . , was unpaved and consisted of a slightly raised central body hardened with gravel and often encased in wood, flanked by (drainage) ditches. With the exception of this road, built in AD 100 and rebuilt in AD 125, in the Netherlands there is little evidence for the existence of roads dating to the first millennium AD.[61]

Rivers, estuaries, and inland waterways such as the Rhine–Meuse system functioned as the main long‑distance transport infrastructure, with rivers offering both opportunities and constraints. They greatly facilitated bulk and high‑volume traffic while making some terrestrial crossings difficult.​ From the sixth century onward, northern branches of the Rhine in the Netherlands regained geographical importance for transportation, setting the stage for intensified Merovingian river trade and North Sea connections.​ By the later seventh to eighth centuries, settlements along the Rhine operated within an extensive exchange network. [62]

Along the northern coast (Frisia sensu lato, from Zeeland/North Holland into Groningen and north‑west Germany), terps and levees framed a string of landing places connected by shallow coastal waterways and tidal channels rather than by continuous built roads. Between the 6th and 8th centuries, Frisian communities developed a maritime commercial system whose backbone was seaborne transport and river mouths, tying the coastal Netherlands to England, the Frankish hinterland and up to the Weser–Elbe region. Environmental studies of the Coastal Frisia area and the north seaways indicate ongoing peatland expansion and marine ingressions in parts of the northern Netherlands between about 700 and 1000 CE, which would have both opened and closed local navigable routes and forced shifts in settlement and paths inland. [63]

From the mid‑seventh century, Dorestad at the Rhine bifurcation of the river branches Kromme Rijn and Lek emerged as a major emporium whose position on a high natural levee along a relatively stable Rhine branch offered sheltered harboring and direct access to both inland and North Sea traffic.​ Dendrochronological and archaeological work shows that Dorestad and earlier coastal sites such as Oegstgeest formed part of a long‑distance network moving timber and other bulk goods along the Rhine–Meuse axis, with riverine transport clearly dominant for heavy cargo.​ New water routes via branches like the Waal and IJssel gradually shifted the main axes of trade in the late eighth to ninth centuries. In the 400–800 CE time window, river corridors in the Meuse–Rhine basin and coastal inlets were already the critical structuring elements of transport routes and communication, far outweighing any surviving terrestrial “road” infrastructure. [64]

Landscape Archaeological Modeling

The ability to reduce this wide range of possible northward migratory paths is strengthened through the use of results from a growing corpus of interdisciplinary archaeological research that identifies general enduring historical route networks or ‘transport corridors‘. These transport corridors can be viewed as a key to understanding large-scale settlement patterns, possible migratory pattterns, and transportation networks.

This general research approach, refered to as ‘landscape and settlement archaeology‘, presents a general theoretical and methodological perspective in archaeology. This research approach utilizes a number of interdisciplinary methodologies that combine historical environmental and archaeological data with the aim to reconstruct probable routes, called movement zones or corridors (see illustration fourteen in the sidebar discussion). [65]

The research from the landscape archaeological perspective provides historical data to uniquely refine and add geographical and historical context to the estimated genetic migratory path derived from the FamilyTreeDNA Globetrekker program. The Globetrekker program provides a statistically smoothed, large‑scale reconstruction of estimated patrilineal migration paths, while landscape archaeological work on connectivity, historically persistent places, and movement corridors can supply the fine‑grained, time‑specific spatial frameworks needed to interpret those migratory paths in real historical landscapes. Used together, they let you move from an abstract Y‑DNA trajectory to concrete hypotheses about which river valleys, trackways, hubs, and settlement zones specific lineages likely used or avoided in particular periods (see sidebar discussion on Landscape and Settlement Archaeology). [66]

The aim of these studies is to bridge gaps between historical and natural science based approaches in studies of the human past through joint research between archaeologists, historians and geoscientists. They are often framed explicitly as contributions to a broader “landscape‑archaeological” and “connectivity/persistence” program for the Dutch delta in the first millennium CE. [67]

Early medieval land and water transport in the Low Countries emerges from this work as dense, highly structured, and remarkably persistent over the first millennium CE, yet constantly re-routed within a dynamic fluvial landscape and shifting political economy. Together these studies show that a limited set of “movement corridors” along levees and navigable channels dominated connectivity from the Roman period into the Early Middle Ages, with most routes continuing in use for centuries and strongly shaping later medieval and even modern infrastructure. [68]

Landscape and Settlement Archaeology

Landscape archaeology can be defined as the interdisciplinary investigation of the long-term relation ship between people and their environment.

Probably the greatest benefit of a landscape – archaeological approach is the way it shifts the focus from a “single-site” perspective to much larger areas that are more closely matched to the physical scale at which human societies operate. Such an approach is in evitably multidisciplinary.

‘Landscape’ within this context is defined at a basic level, being “an area, as perceived by people, whose character is the result of the action and interaction of natural and/or human factors.” [69]

Illustration Fourteen: Flowchart of route-persistence calculations

Click for Larger view | Source: Figure 3 in Rowin J. van Lanen, Bert J. Groenewoudt, Theo Spek & Esther Jansma, Route persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands, Archaeol Anthropol Sci (2018) 10:1037–1052

Their work focuses on how environmental factors (like the changing river network) and cultural factors influenced the orientation and use of persistent water and land travel routes or corridors over time.

These researchers utilize a research approach that integrates layers of archaeological settlement data, geomorphological maps, soil and groundwater data and tree ring data within Geographic Information Systems (GIS) models, to understand the long-distance transport routes and their dynamics during the first millennium CE in the Dutch river delta (see illustration two).

This interdisciplinary approach produces predictive models to reconstruct past movement corridors and assess their persistence through time. Route persistence is studied to explain long‑term settlement foci, land‑use patterns, and “persistent places” in cultural landscapes.

Their research uses high‑resolution palaeogeography, geomorphology, soils, elevation and groundwater reconstructions to map where overland movement was physically feasible in different periods. The landscape is treated as a ‘friction surface’ to predict probable corridors rather than single “roads”. They then combine these models with archaeological proxies (settlements, burial grounds, stray finds, shipwrecks, known roads) and dendroarchaeological timber provenance to validate and refine reconstructed networks and identify long‑distance transport zones. [70]

Modelling from these stduies shows that Roman and early medieval transport relied on intertwined road, track and waterway systems, concentrating movement in relatively narrow route zones that together cover just over ten percent of the Dutch surface but contain roughly three quarters or more of known first‑millennium infrastructural and isolated finds. When Roman and early medieval models are compared, around two‑thirds of the reconstructed corridors persist across the entire first millennium CE, and later early‑modern road networks cluster very strongly on these same long‑lived routes, indicating deep historical stability in movement patterns. [71]

The work underlines that Rhine–Meuse–Scheldt lowland rivers were simultaneously prime highways and major constraints. They offered efficient long‑distance water transport but forced terrestrial routes onto specific levee ridges, crevasse splays and sandy outcrops where flooding risk and groundwater conditions allowed year‑round passage. In delta settings minor avulsions and channel shifts might displace paths locally, yet the overall orientation of corridors and the nodal position of certain confluences and crossings remained stable over centuries. [72]

Collectively, these studies argue that the end of Roman rule did not produce a collapse of connectivity in the Low Country. Instead, networks were reorganized within the same physical framework, with many corridors continuing to channel local, regional and long‑distance movement into the Carolingian period. This long‑term route persistence helps explain the endurance of certain settlement locations and the later emergence of medieval towns along these corridors. It demonstrates that any interpretation of early medieval economic or political change in the lowlands must take seriously the constraining and enabling role of the inherited route system and deltaic landscape.

Globetrekker Migratory Paths and Route Persistence in Landscape Archaeology

The methodogical underpinnings of a cost-efficient migratory path associated with the Globetrekker platform is similar to the concept of route persistence found in these paleogeographical studies of the medieval lowlands. Route persistence is defined as the spatial correlation between route sections across different historical periods. It refers to the long-term use of specific movement corridors or locations, not necessarily continuous use, but rather the tendency for routes to reappear in the same general area over centuries (see sidebar discussion). [73]

This concept ‘route persistence’ is used to investigate the stability of historical ‘transport networks’ and the dynamic interaction between human activity (cultural dynamics) and natural landscape changes over time. Route persistence is studied to explain long‑term settlement locations, land‑use patterns, and “persistent places” in cultural landscapes.

The studies use high‑resolution palaeogeography, geomorphology, soils, elevation and groundwater reconstructions to map where overland movement was physically feasible in different periods, treating the landscape as a ‘friction surface’ to predict probable corridors rather than single “roads”. They then combine these models with archaeological proxies (settlements, burial grounds, stray finds, shipwrecks, known roads) and dendroarchaeological timber provenance to validate and refine reconstructed networks and identify long‑distance transport zones.

Network friction is described as the variable that determines regional accessibility based on local and surrounding landscape factors and that locates transport obstacles and possible movement corridors . Through a network-friction analysis, potential movement corridors are determined. [74]

The Key Aspects of ‘Route Persistence’ in Landscape Archaeological Paleogeographic Studies

Spatial Correlation: Persistence is calculated by determining the degree to which route networks from one era (e.g., Roman period) overlap with those from a later era (e.g., Early Middle Ages) using spatial analysis techniques like GIS (Geographic Information Systems).

“Persistent Places”: The term draws on the concept of “persistent places,” locations that were “never” completely abandoned but survived in collective memory or as logical paths dictated by the physical landscape.

Landscape Influence: In dynamic lowland environments such as the Netherlands, the orientation of routes was highly impacted by natural landscape features. Features like rivers, peat marshes, and levees created natural movement corridors, and routes often persisted in these accessible areas unless major landscape changes (like severe flooding or coastline shifts) occurred. Route persistence is framed as part of regional landscape evolution and human–environment interaction.

Dynamic vs. Fixed Routes: Medieval land routes in lowlands were often unpaved “route zones” rather than fixed, narrow roads. Travelers frequently shifted between parallel tracks within a wider corridor due to seasonal conditions (e.g., moisture, wear and tear), contributing to the idea of a persistent corridor rather than a single, rigid line.

Quantifying Stability: Researchers quantify route persistence (as a percentage of overlap) to understand the relative roles of environmental and cultural factors in shaping the landscape over time. For example, studies have shown a high degree of persistence between Roman and early medieval route networks in the Netherlands, indicating the significant influence of the stable environmental conditions of the time. [75]

Possible Migratory Paths for Ancestors of G-Z6748

An interesting 2015 article by Rowin van Lanen, Menne Kosian, Bert Groenewoudt, and Esther Jansma argues that Roman and early-medieval route networks in the Netherlands can be effectively reconstructed by modelling how key landscape characteristics —especially water, peat, and levees—constrained and channelled the movement on land and water around  100 AD and 800 AD . [76] The study’s main goal is to identify landscape prerequisites for route orientation by using spatial modelling of modern and palaeogeographical data for around these two time perods. The researchers calculate network-friction values for different terrain types and hydrological features, producing maps of likely ‘movement corridors’ and incorporating archaeological data on known routes and sites to be integrated and tested against these models.

In this lowland setting, water bodies, peat zones, and river levees emerge as the dominant landscape characteristics structuring where routes could plausibly run, with substantial contrasts between relatively dry Pleistocene sands [77] and wet coastal and deltaic lowlands. The lower western Netherlands is almost impassable in this time period, implying that inhabitants must have relied predominantly on rivers and streams for transport, while levees and better-drained interfluves acted as preferred terrestrial corridors elsewhere.

As depictived in illustration fifteen below, the research by van Lanen and associates show areas that were inaccessible (in red), moderately accessible (in yellow) and accessible (in green) by land.. Based on this integrated method, geoscientific and archaeological data were used to reconstruct Roman and earlymedieval land and water routes.

Illustration Fifteen: Route networks (land and water) overlaid on network-friction maps of the Netherlands: 100 and 800 CE

Click for Larger View | Source: Figure 1 Route networks (land and water) overlaid on network-friction maps of the Netherlands: AD 100 and 800 in van Lanen, R.J., Groenewoudt, B.J., Spek, T. et al. Route persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands. Archaeol Anthropol Sci 10, 1037–1052 (2018). Route persistence. Modelling and quantifying historical route-network stability

Research models show that Roman and early medieval transport relied on intertwined road, track and waterway systems, concentrating movement in relatively narrow route zones that together cover just over ten percent of the Dutch surface but contain roughly three quarters or more of known first‑millennium infrastructural and isolated archaeological finds. When Roman and early medieval models are compared, around two‑thirds of the reconstructed corridors persist across the entire first millennium AD, and later early‑modern road networks cluster very strongly on these same long‑lived routes, indicating deep historical stability in movement patterns. [78]

The research underlines that Rhine–Meuse–Scheldt lowland rivers were simultaneously prime highways and major constraints. They offered efficient long‑distance water transport but forced terrestrial routes onto specific levee ridges, crevasse splays and sandy outcrops where flooding risk and groundwater conditions allowed year‑round passage. In delta settings this created a ‘braided but structured mosaic’ in which minor avulsions and channel shifts might displace paths locally, yet the overall orientation of corridors and the nodal position of certain confluences and crossings remained stable over centuries. [79]

In another research study, using dendrochronological provenancing of oak timbers (see side bar dicsussion), Jansma, Van Lanen and colleagues reconstruct shifting regional timber flows linking the Netherlands to the German Rhineland, Ardennes–Meuse basin and Scheldt region, and then overlay these with the route models to infer likely long‑distance transport routes. The timber data show changing “frequent‑travel zones” through the first millennium. The Roman‑period flows heavily Rhine‑oriented, with later early medieval phases indicating re‑routed connections and renewed river trade—yet these shifts still track the same core movement corridors identified by the landscape based models.

Esther Jansma , Rowin Van Lanen and Harm Jan Pierik, have provided a short journal contribution that focuses on the Low Countries, specifically the Dutch river delta, and integrates fluvial history with archaeological evidence to reconstruct transport routes over the first millennium CE. The authors identify major river branches and channels that were navigable or strategically important at different times, showing that shifts in discharge and avulsion altered which routes were most suitable for long‑distance movement.​ They highlight that the delta landscape, with its alluvial ridges and natural levees, offered persistent corridors for habitation and movement, even as flooding frequency and local conditions changed.

Dendrochronological Provenancing

Dendrochronological provenancing uses tree-ring analysis (dendrochronology) to determine the geographical origin (provenance) of wood in historical artifacts, buildings, or artworks, not just their age. By matching the unique patterns of wide and narrow rings (influenced by local climate) from an unknown sample to established regional chronologies, researchers identify the source area, often combining ring-width data with chemical analysis (like Sr isotopes) for greater accuracy, especially for timber from complex areas like shipwrecks or Roman structures. [80]

Archaeological remnants of movable wooden objects are well suited for reconstructing past spatial connections because of the following characteristics:

• swift transport and direct application (construction timber);

• river-bound distribution (shipwrecks);

• direct geographical links with the economic hinterland (barrels); and

• high dating precision through dendrochronology.

Absolutely dated dendrochronological time series derived from such objects can be regarded as an integrator of environmental and cultural information, since archaeological wood is the residue of both the site conditions that governed annual tree growth and human activity such as the felling, transport and application of these trees. [81]

Jansma and associates indicate that during this time period several new river branches formed. As a result economic activity shifted from the central and western parts of the delta to the east (see illustrations sixteen and seventeen).

Illustration Sixteen: Long-distance transport routes in the Netherlands during the Roman Period

Click for Larger View | Source: Figure 1 Long-distance transport routes in the Netherlands during the Roman Period. Background: map showing the Roman coast line in Jansma, Esther and Rowin J. Van Lane, and Harm Jan Pierik, Traveling through a River Delta: A Landscape Archaeological Reconstruction of River Development and Long-Distance connections in the Netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39

Illustration Seventeen: Long-distance transport routes in the Netherlands during the Early Middle Ages

Click for Larger View | Source: Figure 2 Long-distance transport routes in the Netherlands during the Early Middle Ages. Background: map showing the early medieval coast line in Jansma, Esther and Rowin J. Van Lane, and Harm Jan Pierik, Traveling through a River Delta: A Landscape Archaeological Reconstruction of River Development and Long-Distance connections in the Netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39

In view of the ‘cost-efficient’ migratory path for the Griff(is)(es)(ith) YDNA line generated by the Globetrekker program (illustration five above), one can add historical context to the migratory path using the results of the study by Jansma and associates (illustration seventeen). It is possible that ancestors of the MCRA of haplogroup G-Z6748 migrated through the changing delta and watershed landscape and utilized what is refered to as the north south corridor to migrate northward via water routes (see illustration eighteen).

Illustration Eighteen: Estimated Migratory Path of G-Z6748 Ancestors Based on Long Distance Alluvial – Water Transport Routes in Early Middle Ages

Click for Larger View | Source: Figure 2 Long-distance transport routes in the Netherlands during the Early Middle Ages, in Esther Jansma, Rowin J. Van Lane, and Harm Jan Pierik, Traveling through a River Delta: A Landscape Archaeological Reconstruction of River Development and Long-Distance connections in the Netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39

The corridor runs roughly north–south across the central–eastern delta (Veluwe/IJssel–Nijmegen–Maas zone), exploiting relatively high, well‑drained ridges and fossil levees that remained passable under rising groundwater and increased flooding. The corridor acted as a backbone between upstream Rhine–Meuse reaches and more northerly and easterly areas, integrating riverine shipping with overland traffic and channelling timber and other bulk goods into and out of the delta.

The north–south corridor is presented as a long‑lived, high‑priority axis of movement that linked the central Rhine–Meuse delta to inland regions and helped maintain regional connectivity despite major changes in river courses and periods of flooding. The corridor exemplifies how terrestrial corridors on elevated levees and higher ground between two rivers stabilize the transport system when fluvial routes shifted or became less reliable.

Landscape archaeological modelling work cited alongside Jansma et al. shows that this north–south zone is one of the most persistent movement corridors: a high proportion of Roman routes in this band were reused or re‑established in early medieval and later networks, and it envelops several later historic towns (e.g. Arnhem, Deventer, Nijmegen). The corridor demonstrates that long‑distance exchange in the first millennium AD depended less on any single river branch and more on a composite system in which enduring overland corridors like this one absorbed and redirected flows when channels avulsed, thereby dampening the impact of environmental instability on migration, trade and communication. [82]

Sources:

Feature Image: The banner depicts the complementary nature of melding the methodologies of the FamilyTreeDNA Globetrekker program with graphic results from the Landscape Archaeological tradition for graphically portraying the migratory path to the ancestor of haplogroup G-6848. The left hand image is a map of the estimated migratory path of ancestors of the Most Recent Ancestor of G-Z6748. The middle images are the scientific details for the estimated birth date for the most recent common ancestor associated with haplogroup G-Z6748. The right hand map depicts a specific migratory path of ancestors of the Most Recent Ancestor of G-Z6748 based on persistent long-distance transport routes in the Netherlands during the Roman period and Early Middle Ages.

[1] See for example:

Cheyette, F. L. , The disappearance of the ancient landscape and the climatic anomaly of the early Middle Ages: A question to be pursued. Early Medieval Europe, 16(2), 2008, 127–165. https://www.academia.edu/395947/The_Disappearance_of_the_Ancient_Landscape_and_the_Climatic_Anomaly_of_the_Early_Middle_Ages_a_Question_to_Be_Pursued

Rowin J van Lanen and Bert Groenewoudt, Counting heads: Post-Roman population decline in the Rhine-Meuse delta (the Netherlands) and the need for more evidence-based reconstructions, in Niall Brady & Claudia Theune, eds, Settlement Change Across Medieval Europe: Old paradigms and New Vistas, , 2019, Ruralia XII, Leiden; Sidestone, 2019, https://www.academia.edu/40523376/Counting_heads_Post_Roman_population_decline_in_the_Rhine_Meuse_delta_the_Netherlands_and_the_need_for_more_evidence_based_reconstructions

Groenewoudt, Bert, Diverging decline. Reconstructing and validating (post-)Roman population trends (AD 0-1000) in the Rhine-Meuse delta (the Netherlands), European Journal of Postclassical Archaeologies (PCA), vol 8, 2018, 189-218, https://www.academia.edu/37462618/Diverging_decline_Reconstructing_and_validating_post_Roman_population_trends_AD_0_1000_in_the_Rhine_Meuse_delta_the_Netherlands_

Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

Pierik, Harm Jan, and Rowin J. van Lanen. “Roman and early-medieval habitation patterns in a delta landscape: The link between settlement elevation and landscape dynamics.” Quaternary International 501 (2019): 379-392. https://www.sciencedirect.com/science/article/abs/pii/S1040618216313453

[2] Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

See the section “The Late-Roman Social and Demographic Collapse” in part seven of this story, The Turbulent Roman Era – The Griff(is)(es)(ith) Y-DNA Phylogenetic Gap Associated with the Meuse and Rhine River Watershed – Part Seven, November 30, 2025, Griffis Family Blog.

de Bruin, Jasper, Connectivity in the south-western part of the Netherlands during the Roman period (AD 0-350), Analecta Praehistorica Leidensia, 43/44, 145 – 157, https://www.academia.edu/30199226/Connectivity_in_the_south_western_part_of_the_Netherlands_during_the_Roman_period_AD_0_350_?email_work_card=view-paper

Kluiving, Sjoerd, Mass migration through soil exhaustion: Transformation of habitation patterns in the southern Netherlands (1000 BC–500AD), Catena (2015), http://dx.doi.org/10.1016/j.catena.2014.12.015 , https://www.academia.edu/20652957/Mass_migration_through_soil_exhaustion_Transformation_of_habitation_patterns_in_the_southern_Netherlands_1000_BC_500AD_?email_work_card=view-paper

[3] Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

[4] FamilyTreeDNA’s “Scientific Details” confidence intervals for haplogroups are derived from their Time to Most Recent Common Ancestor (TMRCA) modeling, which combines the observed genetic variation on each branch with calibrated mutation rates and then summarizes the resulting age uncertainty as statistical intervals. For each haplogroup branch, FTDNA measures its “stem length” in mutations (primarily SNPs, with STRs integrated for very young lineages) and relates this to time using a mutation rate model.

The basic relationship for the mutation rate model is T = D /(2μ), where T is the TMRCA, D is the genetic distance (number of differences between descendants), and μ is the mutation rate. This relationship is statistically ‘refined’ across the whole tree using methods such as linear regression, mean path lengths, maximum likelihood, or relaxed molecular clocks.

Rather than reporting only a single age (“Mean”), FTDNA fits a probability distribution for the branch age that accounts for stochastic mutation processes, rate variation among stems, and tree structure. 

The confidence interval (CI) shown in Scientific Details is the time range that contains a chosen proportion of that distribution (for example, 68%, 95%, or 99% of the total probability), so the narrow, dark band is a higher‑probability, tighter interval and the lighter, wider band is a lower‑probability but more inclusive range.

On a Discover haplogroup page, the Age Estimate section lists a mean age plus one or more CIs; the legend notes that “CI is the Confidence Interval for a given time range and Mean is the average age estimate,” making clear that the interval is a probabilistic range around the model’s best estimate. 

The same underlying methodology is applied across the Y haplotree, with parameters tuned and periodically updated as tree structure, calibration points (including well‑dated historical lineages and selected ancient DNA samples), and rate models are refined. When the algorithm is updated, both the point estimates and their confidence intervals for affected branches can shift.

See: Scientific Details: A Deeper Dive Into Age Estimates, 19, Sep 2022, FTDNA Blog, https://blog.familytreedna.com/tmrca-age-estimates-scientific-details/

[5] The Migration Period (c. 300–700 AD) was a pivotal era in European history, also known as the Barbarian Invasions, characterized by large-scale movements of Germanic, Slavic, and other peoples into and across the declining Western Roman Empire, leading to its collapse, the formation of new post-Roman kingdoms, and shaping modern European cultures and identities. Triggered by factors like Hunnic pressure, climate change, and Roman internal struggles, groups like the Goths, Franks, Vandals, and Slavs settled former Roman territories, transitioning Europe from Late Antiquity into the Early Middle Ages.

Migration Period, Wikipedia, This page was last edited on 19 December 2025, https://en.wikipedia.org/wiki/Migration_Period

[6] After the Roman withdrawal, the main “indigenous” power blocs in the lands of the modern Netherlands are usually identified as Franks in the south, Frisians along the coastal north, and Saxon-identified groups in the east, with all three the result of early medieval ethnogenesis rather than direct continuation of Roman-period tribes. Archaeology and anthropology also stress strong regional continuity from late Roman provincial and “native” communities, so these labels mask a heterogeneous population incorporating Batavian, Cananefatian, Chamavian and other pre-Roman/Roman-period groups.

Political Map of Europe, 651 CE

Click for Larger View | Source: Modified version of map by Richard Ishida, Historical map of Europe in 651 CE, 2016, Wikimedia Commons,https://commons.wikimedia.org/wiki/File:651_CE,_Europe.svg

Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

[7] Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Frisian–Frankish wars, Wikipedia, This page was last edited on 21 December 2025, https://en.wikipedia.org/wiki/Frisian–Frankish_wars

[8] van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences, Volume 99, e14. https://doi.org/10.1017/njg.2020.12

[9] Helama, Samuli and P. Jones, Keith Briffa, Keith, Dark Ages Cold Period: A literature review and directions for future research, February 2017, The Holocene 27(10): 095968361769389, DO – 10.1177/0959683617693898,
https://www.researchgate.net/publication/314130365_Dark_Ages_Cold_Period_A_literature_review_and_directions_for_future_research

[10] Knol, Egge, ‘Living Near the Sea: The Organisation of Frisia in Early Medieval Times’, in Jayne Carroll, Andrew Reynolds, and Barbara Yorke (eds), Power and Place in Europe in the Early Middle Ages, Proceedings of the British Academy (London
, 2019; online edn, British Academy Scholarship Online, 21 May 2020), https://doi.org/10.5871/bacad/9780197266588.003.0017 

Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Terps: Ancient Dwelling Mounds of the North Sea Coast, Ealdlar, https://ealdlar.com/history/terps

Scheurle, Carolyn, Climate development and its effect on the North Sea environment, PhD Dissertation, University of Bremen 2004, https://d-nb.info/975465481/34

[11] The Medieval Climate Anomaly (MCA), also known as the Medieval Warm Period (MWP) or Optimum, was a natural, regional climate event (roughly 900-1300 CE) marked by warmer temperatures

Knol, Egge, ‘Living Near the Sea: The Organisation of Frisia in Early Medieval Times’, in Jayne Carroll, Andrew Reynolds, and Barbara Yorke (eds), Power and Place in Europe in the Early Middle Ages, Proceedings of the British Academy (London
, 2019; online edn, British Academy Scholarship Online, 21 May 2020), https://doi.org/10.5871/bacad/9780197266588.003.0017,

Terps: Ancient Dwelling Mounds of the North Sea Coast, Ealdlar, https://ealdlar.com/history/terps

Weather in History 500 to 750AD, weatherwebdotnet, Accesssed 12 Dec 2025, https://premium.weatherweb.net/weather-in-history-500-to-750ad/

Medieval Warm peiod, Wikipedia, This page was last edited on 29 December 2025, https://en.wikipedia.org/wiki/Medieval_Warm_Period

[12]  Knol E, Ijssennagger N. Palaeogeography and People: Historical Frisians in an archaeological light. In: Hines J, IJssennagger N, eds. Frisians and Their North Sea Neighbours: From the Fifth Century to the Viking Age. Boydell & Brewer; 2017:5-24. https://resolve.cambridge.org/core/books/abs/frisians-and-their-north-sea-neighbours/palaeogeography-and-people-historical-frisians-in-an-archaeological-light/CA5BEE0DAC45640D30C3E40FBDC11C49

Terps: Ancient Dwelling Mounds of the North Sea Coast, Ealdlar, https://ealdlar.com/history/terps

Faber, Hans, A Frontier Known as Watery Mess: the Coast of Flanders, 8 Aug 2021, Frisia Coast Trail Blog, https://frisiacoasttrail.blog/2021/08/08/the-frontier-collectively-known-as-watery-mess-the-coast-flanders/

[13] Knol E, Ijssennagger N. Palaeogeography and People: Historical Frisians in an archaeological light. In: Hines J, IJssennagger N, eds. Frisians and Their North Sea Neighbours: From the Fifth Century to the Viking Age. Boydell & Brewer; 2017:5-24. https://resolve.cambridge.org/core/books/abs/frisians-and-their-north-sea-neighbours/palaeogeography-and-people-historical-frisians-in-an-archaeological-light/CA5BEE0DAC45640D30C3E40FBDC11C49

Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Ogden, Frank, Frankish Law and the Wergeld in the Early Middle Ages, 9 Nov 2020, Ancient, Medieval and Renaissance Studies Program, https://sites.owu.edu/trident/2020/11/09/frankish-law-and-the-wergeld-in-the-early-middle-ages-by-drake-ogden/

Germanic Law, Wikipedia, This page was last edited on 3 December 2025, https://en.wikipedia.org/wiki/Germanic_law

Loveluck C. Context. In: Northwest Europe in the Early Middle Ages, c.AD 600–1150: A Comparative Archaeology. Cambridge University Press; 2013:1-30. https://www.cambridge.org/core/books/abs/northwest-europe-in-the-early-middle-ages-cad-6001150/context/EF9A12D59EB8F07015B489604E75D998

[14] Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

[15] Nicolay, J.A.W. & Nieuwhof, A. “Immobile farmers? The geographical mobility and cultural identity of early medieval Frisians.” Medieval Settlement Research, 2018, 33: 21-31

Knol E, Ijssennagger N. Palaeogeography and People: Historical Frisians in an archaeological light. In: Hines J, IJssennagger N, eds. Frisians and Their North Sea Neighbours: From the Fifth Century to the Viking Age. Boydell & Brewer; 2017:5-24. https://resolve.cambridge.org/core/books/abs/frisians-and-their-north-sea-neighbours/palaeogeography-and-people-historical-frisians-in-an-archaeological-light/CA5BEE0DAC45640D30C3E40FBDC11C49

[16] Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Frisian–Frankish wars, Wikipedia, This page was last edited on 21 December 2025, https://en.wikipedia.org/wiki/Frisian–Frankish_wars

Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences. 2021;100:e11. doi:10.1017/njg.2021.8 , https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/landscape-changes-and-humanlandscape-interaction-during-the-first-millennium-ad-in-the-netherlands/3664067EC3153C9C9B99EBB4871CB770

[17] Harm Jan Pierik, Rowin J. van Lanen, Roman and early-medieval habitation patterns in a delta landscape: The link between settlement elevation and landscape dynamics, Quaternary International, Volume 501, Part B, 2019, Pages 379-392, ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2017.03.010 .
(https://www.sciencedirect.com/science/article/pii/S1040618216313453 )

Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences. 2021;100:e11. doi:10.1017/njg.2021.8 , https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/landscape-changes-and-humanlandscape-interaction-during-the-first-millennium-ad-in-the-netherlands/3664067EC3153C9C9B99EBB4871CB770

Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

[18] Early medieval farms like those near Roomburg and at De Geer in the Rhine delta were small, subsistence focused agricultural settlements with rectangular longhouses, barns, and outbuildings, often clustered in small groups (farmsteads) utilizing the fertile but flood-prone river landscape, with some (like Koudekerk near Roomburg) potentially part of larger estates, showing adaptation from Roman patterns to more localized, community based farming, relying on mixed farming and local resources for survival. 

Key Characteristics:

  • Location and Landscape: Situated on levees or higher ground within the dynamic Rhine-Meuse delta, often bisected by old river channels (crevasses) used for drainage, with settlements like De Geer and Koudekerk showing this adaptation.
  • Farm Layout: Comprised of rectangular longhouses (around 6x20m) for living and animals, plus other structures like storage buildings, hen houses, and possibly pit houses, forming distinct farmsteads.
  • Economic Focus: Primarily subsistence-oriented, focusing on local food production, using basic agricultural technology (animal traction and ploughs were known) and mixed farming (crops, livestock).
  • Community Structure: Often small clusters of farmsteads, possibly forming small villages, with a communal, practical layout rather than planned towns.
  • Relation to Larger Systems: Certain farm sites like Koudekerk near Roomburg might have been part of larger estates (like Holtlant) or functioned as toll points for the Carolingian realm, indicating some integration into broader economies.
  • Continuity and Change: Farms showed continuity from Late Roman times in habitation patterns, but shifted towards smaller, more permanent settlements compared to the earlier dispersed Roman villas, with increased focus on local, intensive farming practices over time. 

See:

W.A. van Es & W.J.H. Verwers, Early Medieval settlements along the Rhine: precursors and contemporaries of Dorestad, Journal of Archaeology in the Low Countries, 2-1 (May 2010), https://jalc.nl/cgi/t/text/get-pdfcfad.pdf?c=jalc%3Bidno%3D0201a01

Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences. 2021;100:e11. doi:10.1017/njg.2021.8 , https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/landscape-changes-and-humanlandscape-interaction-during-the-first-millennium-ad-in-the-netherlands/3664067EC3153C9C9B99EBB4871CB770

[19] See:

Harm Jan Pierik, Rowin J. van Lanen, Roman and early-medieval habitation patterns in a delta landscape: The link between settlement elevation and landscape dynamics, Quaternary International, Volume 501, Part B, 2019, Pages 379-392, ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2017.03.010 .
(https://www.sciencedirect.com/science/article/pii/S1040618216313453 )

Kosian, Menne and Henk Weerts, Rowin Van Lanen, Jaap Evert Abrahamse, The City and the River. The early medieval Emporium (trade centre) of Dorestad; integrating physical geography with archaeological data in changing environments, International Conference on Cultural Heritage and New Technologies, Vienna, 2012, https://chnt.at/wp-content/uploads/eBook_CHNT17_Kosian.pdf

van Dinter, Marieke, Living along the Limes Landscape and settlement in the Lower Rhine Delta during Roman and Early Medieval times, 23 Aug 1970, Utrecht Studies in Earth Sciences 135, https://www.academia.edu/35123998/Living_along_the_Limes_Landscape_and_settlement_in_the_Lower_Rhine_Delta_during_Roman_and_Early_Medieval_times

[20] Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Faber, Hans, The Batwing Doors of Dorestad. A Two-Way Gateway of Trade and Power, 16 Oct 2020, Frisia Coast Trail Blog, https://frisiacoasttrail.blog/2020/10/26/dorestat/

Kosian, Menne and Henk Weerts, Rowin Van Lanen, Jaap Evert Abrahamse, The City and the River. The early medieval Emporium (trade centre) of Dorestad; integrating physical geography with archaeological data in changing environments, International Conference on Cultural Heritage and New Technologies, Vienna, 2012, https://chnt.at/wp-content/uploads/eBook_CHNT17_Kosian.pdf

[21] Faber, Hans, Well, the Thing Is …, 5 Sep 2021, Frisia Trail Coast Blog, https://www.frisiacoasttrail.com/post/the-thing-is

Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Frisian–Frankish wars, Wikipedia, This page was last edited on 21 December 2025, https://en.wikipedia.org/wiki/Frisian–Frankish_wars

W.A. van Es & W.J.H. Verwers, Early Medieval settlements along the Rhine: precursors and contemporaries of Dorestad, Journal of Archaeology in the Low Countries, 2-1 (May 2010), https://jalc.nl/cgi/t/text/get-pdfcfad.pdf?c=jalc%3Bidno%3D0201a01

[22] Netherlands in the Early Middle Ages, Wikipedia, This page was last edited on 2 December 2025, https://en.wikipedia.org/wiki/Netherlands_in_the_Early_Middle_Ages

Kosian, Menne and Henk Weerts, Rowin Van Lanen, Jaap Evert Abrahamse, The City and the River. The early medieval Emporium (trade centre) of Dorestad; integrating physical geography with archaeological data in changing environments, International Conference on Cultural Heritage and New Technologies, Vienna, 2012, https://chnt.at/wp-content/uploads/eBook_CHNT17_Kosian.pdf

Faber, Hans, The Batwing Doors of Dorestad. A Two-Way Gateway of Trade and Power, 16 Oct 2020, Frisia Coast Trail Blog, https://frisiacoasttrail.blog/2020/10/26/dorestat/

van Popta, Y.T., Westerdahl, C.L. and Duncan, B.G. (2019), Maritime Culture in the Netherlands: accessing the late medieval maritime cultural landscapes of the north-eastern Zuiderzee. International Journal of Nautical Archaeology, 48: 172-188. https://doi.org/10.1111/1095-9270.12333

[23] Pierik HJ. , Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

P.C. Vos & E. Knol, Holocene landscape reconstruction of the Wadden Sea area between Marsdiep and Weser, Netherlands Journal of Geosciences — Geologie en Mijnbouw ,94 – 2, 157–183, 2015, https://www.waddenacademie.nl/fileadmin/inhoud/pdf/03-Thema_s/Geowetenschap/vos_knol.pdf

Kaspers, Angelique, From sherds to settlement patterns – new insights into the habitation history of the coastal area of the northern Netherlands during the Merovingian and Carolingian periods (AD 400-900) based on field surveys and older collections research, Settlement and Coastal Research in the Southern North Sea Region (SCN), 46, 115-189, Rahden/Westf, 2023, https://www.academia.edu/123867114/From_sherds_to_settlement_patterns_new_insights_into_the_habitation_history_of_the_coastal_area_of_the_northern_Netherlands_during_the_Merovingian_and_Carolingian_periods_AD_400_900_based_on_field_surveys_and_older_collections_research

Hines, John, and Nelleke IJssennagger-van der Pluijm, editors. Frisians of the Early Middle Ages. NED-New edition, Boydell & Brewer, 2021. JSTOR, https://doi.org/10.2307/j.ctv199tj69

H.J. Pierika, K.M. Cohena,, P.C. Vos, A.J.F. van der Spekd, E. Stouthamer, Late Holocene coastal-plain evolution of the Netherlands: the role of natural preconditions in human-induced sea ingressions, Proceedings of the Geologists’ Association 128 (2017) 180–197, https://dspace.library.uu.nl/bitstream/handle/1874/348622/late.pdf

[24] Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

[25] Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

[26] Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

[27] The GlobeTrekker computer program integrates genetic, geological, and anthropological evidence to reconstruct global paternal migration histories. It visualizes how each YDNA haplogroup, down to an individual’s terminal SNP, fits within humanity’s evolving distribution through Ice Age sea-levels, population dispersals, and environmental corridors.

In FamilyTreeDNA’s GlobeTrekker system, corridor paths represent least-cost migration corridors (LCCs) — probabilistic zones indicating the most likely routes Y-DNA lineages followed between ancestral haplogroup locations.  GlobeTrekker identifies each ancestral haplogroup’s probable position and then connects them using these cost-efficient routes that consider:

  • Past topography and sea levels, including exposed Ice Age land bridges and glacial boundaries.;
  • Slope steepness (to avoid rugged terrain);
  • Distance to land (favoring coastlines); and, if appropriate,
  • Ocean current direction and strength (penalizing movement against currents).

The corridor paths serve as migration ‘confidence envelopes’, capturing the uncertainty inherent in reconstructing prehistorical movement based on environmental and genetic data. They combine data from Big Y testers, ancient DNA, and ecogeographic models to show how and where paternal lineages likely spread worldwide over tens of millennia. The visible “corridor bands” in GlobeTrekker are explicitly tied to likelihood percentages, but the single thin line itself is just the minimum-cost path within those bands rather than a separately quantified probability.

See:

Vilar, Miguel, Join us on this extraordinary voyage through time and genetics, where every strand of DNA is a thread in the tapestry of human history, FTDNA Blog, 26 Sep 2023, https://blog.familytreedna.com/globetrekker-history/

Maier, Paul, Globetrekker, Part 2: Advancing the Science of Phylogeography, 15 Aug, 2023, https://blog.familytreedna.com/globetrekker-analysis/

[28] Texel, Wikipedia, This page was last edited on 26 November 2025, https://en.wikipedia.org/wiki/Texel

List of Islands of the Netherlands, Wikipedia, This page was last edited on 1 August 2025,  https://en.wikipedia.org/wiki/List_of_islands_of_the_Netherlands

[29] The creation of Texel, Texel, https://www.texel.net/en/about-texel/history/the-creation-of-texel/

Texel, Wikipedia, This page was last edited on 26 November 2025, https://en.wikipedia.org/wiki/Texel

[30] Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8 ; for access to the article: https://www.researchgate.net/publication/351236350_Landscape_changes_and_human-landscape_interaction_during_the_first_millennium_AD_in_the_Netherlands

[31] “Geomorphological” relates to geomorphology, the scientific study of landforms, their origins, evolution, and the processes (like erosion, weathering, tectonics) that shape them, aiming to understand why landscapes look the way they do and how they change over time due to natural forces and human activity. It’s a core part of physical geography, using field observation, modeling, and technology to analyze features from river valleys and coastlines to mountain ranges. 

Geomophology, Wikipedia, This page was last edited on 5 November 2025, https://en.wikipedia.org/wiki/Geomorphology

[32] Jansma, Esther, Hydrological disasters in the NW-European Lowlands during the first millennium AD: a dendrochronological reconstruction, Netherlands Journal of Geosciences, Volume 99, e11. https://doi.org/ 10.1017/njg.2020.10 https://www.academia.edu/44188764/Jansma_E_2020_Hydrological_disasters_in_the_NW_European_Lowlands_during_the_first_millennium_AD_a_dendrochronological_reconstruction

For another study that focuses on the Rhine River flooding, see:

Toonen, W.H.J., Donders, T.H., Van der Meulen, B., Cohen, K.M. and Prins, M.A. 2013. A composite Holocene palaeoflood chronology of the Lower Rhine. In W.H.J Toonen (ed.), A Holocene Flood Record of the Lower Rhine, Utrecht Studies in Earth Sciences 41: 137–150.

[33] Jansma, Esther, Hydrological disasters in the NW-European Lowlands during the first millennium AD: a dendrochronological reconstruction, Netherlands Journal of Geosciences, Volume 99, e11. https://doi.org/ 10.1017/njg.2020.10 https://www.academia.edu/44188764/Jansma_E_2020_Hydrological_disasters_in_the_NW_European_Lowlands_during_the_first_millennium_AD_a_dendrochronological_reconstruction

[34] Frisia, Wikipedia, This page was last edited on 29 November 2025, https://en.wikipedia.org/wiki/Frisia

Flierman R. Mirror Histories: Frisians and Saxons from the First to the Ninth Century AD. In: Hines J, IJssennagger-van der Pluijm N, eds. Frisians of the Early Middle Ages. Studies in Historical Archaeoethnology. Boydell & Brewer; 2021:223-248, https://research-portal.uu.nl/ws/portalfiles/portal/232053797/mirror-histories-frisians-and-saxons-from-the-first-to-the-ninth-century-ad.pdf

[35] Terp, Wikipedia, This page was last edited on 17 August 2025, https://en.wikipedia.org/wiki/Terp

A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
(https://www.sciencedirect.com/science/article/pii/S0964569118307348 )

[36] Terps: Ancient Dwelling Mounds of the North Sea Coast, no date, Ealdlar, https://ealdlar.com/history/terps

See also:

Nicolay, J.A.W. & Nieuwhof, A. (2018). “Immobile farmers? The geographical mobility and cultural identity of early medieval Frisians.” Medieval Settlement Research, 33: 21-31,

Nieuwhof, Annet, Discontinuity in the Northern-Netherlands coastal area at the end of the Roman Period, in: Transformations in North-Western Europe (AD 300-1000) Proceedings of the 60th Sachsen symposion 19.-23. September 2009 Maastricht”, Hanover: Die Publishing Company, 2011, https://www.academia.edu/1449600/Discontinuity_in_the_Northern_Netherlands_coastal_area_at_the_end_of_the_Roman_Period

[37] Progradation is the geological process where a landform, like a river delta, beach, or alluvial fan, builds outward into a body of water (seaward or basinward) due to continuous sediment accumulation, causing the shoreline to advance, often linked to sea-level fall or high sediment supply.

Progradation, Wikipeida, This page was last edited on 14 August 2025, https://en.wikipedia.org/wiki/Progradation

[38] Terps: Ancient Dwelling Mounds of the North Sea Coast, Ealdlar, https://ealdlar.com/history/terps

A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
(https://www.sciencedirect.com/science/article/pii/S0964569118307348 )

J. Bazelmans Joseph and D. Meier, A. Nieuwhof, T. Spekc, P. Vos, Understanding the cultural historical value of the Wadden Sea region. The co-evolution of environment and society in the Wadden Sea area in the Holocene up until early modern times (11,700 BCe1800 AD): An outline,  Ocean & Coastal Management 68 (2012) 114-126, https://www.waddenacademie.nl/fileadmin/inhoud/pdf/04-bibliotheek/Themanummer_OCMA/11_Understanding_the_cultural_historical_value_of_the_Wadden_Sea_region_OCMA.pdf

Knol, Egge, ‘Living Near the Sea: The Organisation of Frisia in Early Medieval Times’, in Jayne Carroll, Andrew Reynolds, and Barbara Yorke (eds), Power and Place in Europe in the Early Middle Ages, Proceedings of the British Academy (London
, 2019; online edn, British Academy Scholarship Online, 21 May 2020), https://doi.org/10.5871/bacad/9780197266588.003.0017

[39] A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
(https://www.sciencedirect.com/science/article/pii/S0964569118307348 )

Reiß, A., Hadler, H., Wilken, D., Majchczack, B. S., Blankenfeldt, R., Bäumler, S., Ickerodt, U., Klooß, S., Willershäuser, T., Rabbel, W., and Vött, A.: The Trendermarsch sunken in the Wadden Sea (North Frisia, Germany) – reconstructing a drowned medieval cultural landscape with geoarchaeological and geophysical investigations, E&G Quaternary Sci. J., 74, 37–57, https://doi.org/10.5194/egqsj-74-37-2025 , 2025. See additional link https://egqsj.copernicus.org/articles/74/37/2025/

Bakker, M., The Nature and Dynamics of Pre-Roman Iron Age and Roman Iron Age Reclamation Settlements in the (Former) Peat and Clay-On-Peat Area of Friesland (The Netherlands). Journal of Wetland Archaeology22(1–2), 2022, 7–29. https://doi.org/10.1080/14732971.2022.2061783

[40] A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
(https://www.sciencedirect.com/science/article/pii/S0964569118307348 )

See also:

Faber, Hans, Between Leffinge and Misthusum—Understanding the Basics of Terps, 29 Sep 2024, Frisia Coast Trail, https://www.frisiacoasttrail.com/post/between-leffinge-and-misthusum-understanding-the-basics-of-terps

Knol, Egge, ‘Living Near the Sea: The Organisation of Frisia in Early Medieval Times’, in Jayne Carroll, Andrew Reynolds, and Barbara Yorke (eds), Power and Place in Europe in the Early Middle Ages, Proceedings of the British Academy (London
, 2019; online edn, British Academy Scholarship Online, 21 May 2020), https://doi.org/10.5871/bacad/9780197266588.003.0017

Ellen McManus, Janet Montgomery, Jane Evans, Angela Lamb, Rhea Brettelld, Johan Jelsmae, ‘To the land or to the sea’: diet and mobility in early medieval Frisia, https://nora.nerc.ac.uk/id/eprint/504785/1/McManus%20et%20al%20-%20Oosterbeintum_final.pdf

Bakker, M., The Nature and Dynamics of Pre-Roman Iron Age and Roman Iron Age Reclamation Settlements in the (Former) Peat and Clay-On-Peat Area of Friesland (The Netherlands). Journal of Wetland Archaeology22(1–2), 2022, 7–29. https://doi.org/10.1080/14732971.2022.2061783

[41] Knol, Egge, ‘Living Near the Sea: The Organisation of Frisia in Early Medieval Times’, in Jayne Carroll, Andrew Reynolds, and Barbara Yorke (eds), Power and Place in Europe in the Early Middle Ages, Proceedings of the British Academy (London
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Ellen McManus, Janet Montgomery, Jane Evans, Angela Lamb, Rhea Brettelld, Johan Jelsmae, ‘To the land or to the sea’: diet and mobility in early medieval Frisia, https://nora.nerc.ac.uk/id/eprint/504785/1/McManus%20et%20al%20-%20Oosterbeintum_final.pdf

Terps: Ancient Dwelling Mounds of the North Sea Coast, Ealdlar, https://ealdlar.com/history/terps

Bakker, M., The Nature and Dynamics of Pre-Roman Iron Age and Roman Iron Age Reclamation Settlements in the (Former) Peat and Clay-On-Peat Area of Friesland (The Netherlands). Journal of Wetland Archaeology22(1–2), 2022, 7–29. https://doi.org/10.1080/14732971.2022.2061783

Reiß, A., Hadler, H., Wilken, D., Majchczack, B. S., Blankenfeldt, R., Bäumler, S., Ickerodt, U., Klooß, S., Willershäuser, T., Rabbel, W., and Vött, A.: The Trendermarsch sunken in the Wadden Sea (North Frisia, Germany) – reconstructing a drowned medieval cultural landscape with geoarchaeological and geophysical investigations, E&G Quaternary Sci. J., 74, 37–57, https://doi.org/10.5194/egqsj-74-37-2025 , 2025. See additional link https://egqsj.copernicus.org/articles/74/37/2025/

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Nieuwhof, Annet, Creating a home. Ritual practice in a terp settlement in the northern Netherlands, Beiträge zur Ur- und Frühgeschichte Mitteleuropas 95, 95 – 109, https://www.academia.edu/44905134/Creating_a_home_Ritual_practice_in_a_terp_settlement_in_the_northern_Netherlands

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Faber, Hans, Between Leffinge and Misthusum—Understanding the Basics of Terps, 29 Sep 2024,Frisia Coast Trail Blog, https://www.frisiacoasttrail.com/post/between-leffinge-and-misthusum-understanding-the-basics-of-terps

Niewhof, Annet, Ezinge revisited. The ancient roots of a terp settlement: Volume 1: Excavation – Environment and Economy – Catalogue of Plans and Finds, Groningen Archaeological Studies, 37, University of Groningen/Groningen Institute of Archaeology and Barkhuis Publishing, Oct 2020, https://research.rug.nl/en/publications/ezinge-revisited-the-ancient-roots-of-a-terp-settlement-volume-1-/

[45] Carroll, Jayne, Andrew Reynolds and Barbara York, eds. Power and Place in Europe in the Early Middle Ages. Vol. 224. The British Academy, 2019. http://www.jstor.org/stable/jj.31732145

Nieuwhof, Annet, Anglo-Saxon immigration or continuity? Ezinge and the coastal area of the northern Netherlands in the Migration Period, Journal of Archaeology in the Low Countries 4-2 (April 2013), https://jalc.nl/cgi/t/text/text-idxaa82.html

Angelique Kaspers, Gilles J. de Langen and Johan A. W. Nicolay, From sherds to settlement patterns – new insights into the habitation history of the coastal area of the northern Netherlands during the Merovingian and Carolingian periods (AD 400-900) based on field surveys and older collections research, Settlement and Coastal Research in the Southern North Sea Region (SCN), 46, 115-189, Rahden/Westf. 2023, https://www.academia.edu/123867114/From_sherds_to_settlement_patterns_new_insights_into_the_habitation_history_of_the_coastal_area_of_the_northern_Netherlands_during_the_Merovingian_and_Carolingian_periods_AD_400_900_based_on_field_surveys_and_older_collections_research

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[46] In archaeology, stratigraphy is the study of layers (strata) of soil and debris that build up over time, allowing archaeologists to establish a chronological sequence of events at a site. It relies on the Law of Superposition, which states that deeper layers are older than those above them, helping reconstruct the site’s history, understand past activities, and date artifacts relative to each other, much like layers in a cake or lasagna.

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Kaspers, Angelique and Gilles J. de Langen and Johan A. W. Nicolay, From sherds to settlement patterns – new insights into the habitation history of the coastal area of the northern Netherlands during the Merovingian and Carolingian periods (AD 400-900) based on field surveys and older collections research, Settlement and Coastal Research in the Southern North Sea Region (SCN), 46, 115-189, Rahden/Westf. 2023, https://www.academia.edu/123867114/From_sherds_to_settlement_patterns_new_insights_into_the_habitation_history_of_the_coastal_area_of_the_northern_Netherlands_during_the_Merovingian_and_Carolingian_periods_AD_400_900_based_on_field_surveys_and_older_collections_research

Nieuwhof A. The Frisians and their Pottery: Social Relations before and after the Fourth Century AD. In: Hines J, IJssennagger-van der Pluijm N, eds. Frisians of the Early Middle Ages. Studies in Historical Archaeoethnology. Boydell & Brewer; 2021:45-78, https://resolve.cambridge.org/core/books/abs/frisians-of-the-early-middle-ages/frisians-and-their-pottery-social-relations-before-and-after-the-fourth-century-ad/49D9B6667528E049632B1978BCB811F5

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Gerrets, Danny, Continuous development in house-type and settlement structure in Drenthe, Ruralia Památky archeologické – Supplementum 5, Praha 1996, http://ruralia2.ff.cuni.cz/wp-content/uploads/2018/04/Danny-Gerrets.pdf

Nieuwhof, Annet, Ezinge revisited. The ancient roots of a terp settlement: Volume 1: Excavation – Environment and Economy – Catalogue of Plans and Finds, Gronongen: Univesrity of Groningen / Groningen Institute of Archaeology and Barkhuis Publishing, 2020, https://research.rug.nl/en/publications/ezinge-revisited-the-ancient-roots-of-a-terp-settlement-volume-1-/

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[49] Kaspers, Angelique and Gilles J. de Langen and Johan A. W. Nicolay, From sherds to settlement patterns – new insights into the habitation history of the coastal area of the northern Netherlands during the Merovingian and Carolingian periods (AD 400-900) based on field surveys and older collections research, Settlement and Coastal Research in the Southern North Sea Region (SCN), 46, 115-189, Rahden/Westf. 2023,

Boeles en Van Giffen Speech or profile paper about mounds, Education, Vereniging voor Terpenonderzoek, https://terpenonderzoek.nl/education/?lang=en

[50] Knol, Egge, ‘Living Near the Sea: The Organisation of Frisia in Early Medieval Times’, in Jayne Carroll, Andrew Reynolds, and Barbara Yorke (eds), Power and Place in Europe in the Early Middle Ages, Proceedings of the British Academy (London,  2019; online edn, British Academy Scholarship Online, 21 May 2020), https://doi.org/10.5871/bacad/9780197266588.003.0017

Carroll, Jayne, Andrew Reynolds amd Barbara Yorke, eds. Power and Place in Europe in the Early Middle Ages. Vol. 224. The British Academy, 2019. http://www.jstor.org/stable/jj.31732145

[51] van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
(https://www.sciencedirect.com/science/article/pii/S0964569118307348 )

Knol, Egge, ‘Living Near the Sea: The Organisation of Frisia in Early Medieval Times’, in Jayne Carroll, Andrew Reynolds, and Barbara Yorke (eds), Power and Place in Europe in the Early Middle Ages, Proceedings of the British Academy (London
, 2019; online edn, British Academy Scholarship Online, 21 May 2020), https://doi.org/10.5871/bacad/9780197266588.003.0017

[52] Bakker, M. , The Nature and Dynamics of Pre-Roman Iron Age and Roman Iron Age Reclamation Settlements in the (Former) Peat and Clay-On-Peat Area of Friesland (The Netherlands). Journal of Wetland Archaeology22(1–2), 2022, 7–29. https://doi.org/10.1080/14732971.2022.2061783

van Popta, Y. T., Cohen, K. M., Vos, P. C., & Spek, Th. (2020). Reconstructing medieval eroded landscapes of the north-eastern Zuyder Zee (the Netherlands): a refined palaeogeographical time series of the Noordoostpolder between a.d. 1100 and 1400. Landscape History41(2), 27–56. https://doi.org/10.1080/01433768.2020.1835180

A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
(https://www.sciencedirect.com/science/article/pii/S0964569118307348 )

[53] D. Postma , Salt Marsh Architecture, Catalogue, technology and typological development of early medieval turf buildings in the northern coastal area of the Netherlands, Masters Thesis University of Groningen, 2010, https://www.waddenacademie.nl/fileadmin/inhoud/pdf/06-wadweten/Scripties/D._Postma_-_scriptieverkl.pdf

[54] A. Nieuwhof, M. Bakker, E. Knol, G.J. de Langen, J.A.W. Nicolay, D. Postma, M. Schepers, T.W. Varwijk, P.C. Vos, Adapting to the sea: Human habitation in the coastal area of the northern Netherlands before medieval dike building, Ocean & Coastal Management, Volume 173, 2019, Pages 77-89, ISSN 0964-5691, https://doi.org/10.1016/j.ocecoaman.2019.02.014 .
(https://www.sciencedirect.com/science/article/pii/S0964569118307348 )

Bakker, M. , The Nature and Dynamics of Pre-Roman Iron Age and Roman Iron Age Reclamation Settlements in the (Former) Peat and Clay-On-Peat Area of Friesland (The Netherlands). Journal of Wetland Archaeology22(1–2), 2022, 7–29. https://doi.org/10.1080/14732971.2022.2061783

[55] van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

Van Lanen, R.J., Jansma, E., Van Doesburg, J., Groenewoudt, B.J., Roman and early-medieval long-distance transport routes in north-western Europe: modelling frequent-travel zones using a dendroarchaeological approach. J. Archaeol. Sci. 73, 120e137. 2016, https://www.sciencedirect.com/science/article/abs/pii/S0305440316300978

W.A. van Es & W.J.H. Verwers, Early Medieval settlements along the Rhine: precursors and contemporaries of Dorestad, Journal of Archaeology in the Low Countries, 2-1 (May 2010), https://jalc.nl/cgi/t/text/get-pdfcfad.pdf?c=jalc%3Bidno%3D0201a01

[56] van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

[57] Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

van Lanen, Rowin J. and Esther Jansma, Jan van Doesburg, Bert J. Groenewoudt, Roman and early-medieval long-distance transport routes in north-western Europe: Modelling frequent-travel zones using a dendroarchaeological approach , Journal of Archaeological Science, 73, Sep 2016, 120- 137, https://www.sciencedirect.com/science/article/abs/pii/S0305440316300978

R.J. Lanen, M. Kosian, B.J. Groenewoudt, E. Jansma, Geoarchaeology: An international Journal (30), 200-222(2015): Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands pp. 200 – 222, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands

van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

[58] van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

[59] van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

Jansma, Esther and Rowin J. Van Lanen and Harm Jan Pierk, Traveling Through a River Delta: A Landscape Archaeogical Reconstruction of River Development and Long Distance Connections in the netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39, https://www.academia.edu/35074210/Jansma_E_R_J_Van_Lanen_and_H_J_Pierik_2017_Travelling_through_a_river_delta_a_landscape_archaeological_reconstruction_of_river_development_and_long_distance_connections_in_the_Netherlands_during_the_first_millennium_AD_Medieval_Settlement_Research_32_35_39

Van Lanen, R.J., Kosian, M.C., Groenewoudt, B.J., Spek, T., Jansma, E., 2016b. Route persistence. Modelling and quantifying historical route-network stability during the last two millennia: a case study from The Netherlands. Archaeol. Anthropol. Sciences 1e16. https://www.researchgate.net/publication/310757823_Route_persistence_Modelling_and_quantifying_historical_route-network_stability_from_the_Roman_period_to_early-modern_times_AD_100-1600_a_case_study_from_the_Netherlands

[60] Rowin J. van Lanen, Rowin J. and Harm Jan Pierik, Calculating connectivity patterns in delta landscapes: Modelling Roman and early-medieval route networks and their stability in dynamic lowlands, Quaternary International, Volume 501, Part B, 2019, Pages 393-412,
ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2017.03.009 .
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[61] Quote:

Jansma, Esther and Rowin J. Van Lanen, and Harm Jan Pierik, Traveling through a River Delta: A Landscape Archaeological Reconstruction of River Development and Long-Distance connections in the Netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39, https://www.academia.edu/35074210/Jansma_E_R_J_Van_Lanen_and_H_J_Pierik_2017_Travelling_through_a_river_delta_a_landscape_archaeological_reconstruction_of_river_development_and_long_distance_connections_in_the_Netherlands_during_the_first_millennium_AD_Medieval_Settlement_Research_32_35_39

The quote references the “old Rhine’. “In ancient times, it was the lower part of the main River Rhine, which forked at the Betuwe into a northern branch, the Rhine, and a southern branch, the Waal. The Oude Rijn was then much wider than it is now, and tidal. During the Roman occupation, the river formed part of the northern border of the Empire. In medieval times, the River Lek became the main outlet for the Rhine, and the Oude Rijn silted up.

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[62] W.A. van Es & W.J.H. Verwers, Early Medieval settlements along the Rhine: precursors and contemporaries of Dorestad, Journal of Archaeology in the Low Countries, 2-1 (May 2010), https://jalc.nl/cgi/t/text/get-pdfcfad.pdf?c=jalc%3Bidno%3D0201a01

van Lanen, Rowin J. and Esther Jansma, Jan van Doesburg, Bert J. Groenewoudt, Roman and early-medieval long-distance transport routes in north-western Europe: Modelling frequent-travel zones using a dendroarchaeological approach , Journal of Archaeological Science, 73, Sep 2016, 120- 137, https://www.sciencedirect.com/science/article/abs/pii/S0305440316300978

van Lanen, Rowin J.  and Jaap Evert Abrahamse, The City and the River. The early medieval Emporium (trade centre) of Dorestad; integrating physical geography with archaeological data in changing environments, in Wolfgang Börner, Susanne Uhlirz, Proceedings of the 17th International Conference on Cultural Heritage and New Technologies 2012 , https://www.academia.edu/5851685/The_City_and_the_River_The_early_medieval_Emporium_trade_centre_of_Dorestad_integrating_physical_geography_with_archaeological_data_in_changing_environments

[63] Hines, John, and Nelleke IJssennagger, eds. Frisians and Their North Sea Neighbours: From the Fifth Century to the Viking Age. NED-New edition. Boydell & Brewer, 2017. http://www.jstor.org/stable/10.7722/j.ctt1t6p55t

Pierik, Ham Jan, Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

Flierman, Robert, Mirror Histories: Frisians and Saxons from the First to the Ninth century AD, 223 – 247, Chapter 8 in: Hines J, IJssennagger-van der Pluijm N, eds. Frisians of the Early Middle Ages. Studies in Historical Archaeoethnology. Boydell & Brewer, Published online by Cambridge University Press:  11 June 2021, https://resolve.cambridge.org/core/books/abs/frisians-of-the-early-middle-ages/mirror-histories-frisians-and-saxons-from-the-first-to-the-ninth-century-ad/86AE8F89C42C13A6B84B63D3D06405A6

Tys, Dries, ‘Maritime and River Traders, Landing Places, and Emporia Ports in the Merovingian Period in and Around the Low Countries’, in Bonnie Effros, and Isabel Moreira (eds), The Oxford Handbook of the Merovingian World (2020; online edn, Oxford Academic, 8 Oct. 2020), https://doi.org/10.1093/oxfordhb/9780190234188.013.26

[64] Dorestad, Wikipedia, This page was last edited on 16 December 2024, https://en.wikipedia.org/wiki/Dorestad

van Lanen, Rowin J.  and Jaap Evert Abrahamse, The City and the River. The early medieval Emporium (trade centre) of Dorestad; integrating physical geography with archaeological data in changing environments, in Wolfgang Börner, Susanne Uhlirz, Proceedings of the 17th International Conference on Cultural Heritage and New Technologies 2012 , https://www.academia.edu/5851685/The_City_and_the_River_The_early_medieval_Emporium_trade_centre_of_Dorestad_integrating_physical_geography_with_archaeological_data_in_changing_environments

van Lanen, Rowin J. and Esther Jansma, Jan van Doesburg, Bert J. Groenewoudt, Roman and early-medieval long-distance transport routes in north-western Europe: Modelling frequent-travel zones using a dendroarchaeological approach , Journal of Archaeological Science, 73, Sep 2016, 120- 137, https://www.sciencedirect.com/science/article/abs/pii/S0305440316300978

W.A. van Es & W.J.H. Verwers, Early Medieval settlements along the Rhine: precursors and contemporaries of Dorestad, Journal of Archaeology in the Low Countries, 2-1 (May 2010), https://jalc.nl/cgi/t/text/get-pdfcfad.pdf?c=jalc%3Bidno%3D0201a01

[65] See for example:

Rowin J. van Lanen, Rowin J. and Harm Jan Pierik, Calculating connectivity patterns in delta landscapes: Modelling Roman and early-medieval route networks and their stability in dynamic lowlands, Quaternary International, Volume 501, Part B, 2019, Pages 393-412,
ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2017.03.009 .
(https://www.sciencedirect.com/science/article/pii/S1040618216313519 )

van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

Jansma, Esther and Rowin J. Van Lanen and Harm Jan Pierk, Traveling Through a River Delta: A Landscape Archaeogical Reconstruction of River Development and Long Distance Connections in the Netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39, https://www.academia.edu/35074210/Jansma_E_R_J_Van_Lanen_and_H_J_Pierik_2017_Travelling_through_a_river_delta_a_landscape_archaeological_reconstruction_of_river_development_and_long_distance_connections_in_the_Netherlands_during_the_first_millennium_AD_Medieval_Settlement_Research_32_35_39

Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

Roymans, N., & Gerritsen, F.A. (2002). Landscape, ecology and mentalities: a long-term perspective on developments in the Meuse-Demer-Scheldt region. Proceedings of the Prehistoric Society, 68, 257–287

[66] The idea of couching the results of mapping specific ‘cost-efficient’ migration routes of specific YDNA lineages created by Globetrekker in the context of land archaeological models of general land and water transport route zones, movemement paths and persistence places is similar to a suggested research approach for landscape genomics and genetics.

An editorial by Samuel Cushman and other research associates frames landscape genomics as an expansion of landscape genetics made possible by next generation sequencing (NGS) methods, and concludes that progress hinges on integrating large genomic datasets with spatial modeling and experimental work in hypothesis‑driven, collaborative projects. The article’s main goal is to outline conceptual and practical steps for moving from traditional landscape genetics (e.g. tens of microsatellites, population‑level sampling) to landscape genomics (e.g. thousands of genome‑wide markers from NGS) in a spatially explicit framework.

See: Cushman SA, Shirk AJ, Howe GT, Murphy MA, Dyer RJ, Joost S. Editorial: The Least Cost Path From Landscape Genetics to Landscape Genomics: Challenges and Opportunities to Explore NGS Data in a Spatially Explicit Context. Front Genet. 2018 Jun 19;9:215. doi: 10.3389/fgene.2018.00215. PMID: 29971091; PMCID: PMC6018102 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC6018102/

[67] Jansma et. al. and Van Lanen et. al. discuss the interdisciplinary nature of this research:

Jansma, E., Gouw-Bouman, M., Van Lanen, R., Pierik, H.J., Cohen, K., Groenewoudt, B., Hoek, W., Stouthamer, E., & Middelkoop, H., The dark age of the lowlands in an interdisciplinary light: People, landscape and climate in the Netherlands between AD 300–1000. European Journal of Post-Classical Archaeologies (PCA 4), 2014, 471–476. ISSN: 2039–7895, https://www.researchgate.net/publication/265096511_The_Dark_Age_of_the_Lowlands_in_an_interdisciplinary_light_people_landscape_and_climate_in_The_Netherlands_between_AD_300_and_1000

Van Lanen , Rowin J. and Esther Jansma, and Menne Kosian, Finding a way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An international Journal (30), 200-222. https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

[68] See for example:

Jansma, E., Gouw-Bouman, M., Van Lanen, R., Pierik, H.J., Cohen, K., Groenewoudt, B., Hoek, W., Stouthamer, E., & Middelkoop, H., The dark age of the lowlands in an interdisciplinary light: People, landscape and climate in the Netherlands between AD 300–1000. European Journal of Post-Classical Archaeologies (PCA 4), 2014, 471–476. ISSN: 2039–7895, https://www.researchgate.net/publication/265096511_The_Dark_Age_of_the_Lowlands_in_an_interdisciplinary_light_people_landscape_and_climate_in_The_Netherlands_between_AD_300_and_1000

Jansma, Esther and Rowin J. Van Lanen and Harm Jan Pierk, Traveling Through a River Delta: A Landscape Archaeogical Reconstruction of River Development and Long Distance Connections in the Netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39, https://www.academia.edu/35074210/Jansma_E_R_J_Van_Lanen_and_H_J_Pierik_2017_Travelling_through_a_river_delta_a_landscape_archaeological_reconstruction_of_river_development_and_long_distance_connections_in_the_Netherlands_during_the_first_millennium_AD_Medieval_Settlement_Research_32_35_39

Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8

Roymans, N., & Gerritsen, F.A. (2002). Landscape, ecology and mentalities: a long-term perspective on developments in the Meuse-Demer-Scheldt region. Proceedings of the Prehistoric Society, 68, 257–287

Stouthamer, E., & Berendsen, H.J.A., Factors controlling the Holocene avulsion history of the Rhine–Meuse Delta (the Netherlands). Journal of Sedimentary Research, 70(5), 2000, 1051–1064.

van Dinter, Marieke and Kim M. Cohen, Wim Z. Hoek, Esther Stouthamer, Esther Jansma , Hans Middelkoop, Late Holocene lowland fluvial archives and geoarchaeology: Utrecht’s case study of Rhine river abandonment under Roman and Medieval settlement, Quaternary Science Reviews (2017) 1-39, https://www.academia.edu/31299345/Late_Holocene_lowland_fluvial_archives_and_geoarchaeology_Utrechts_case_study_of_Rhine_river_abandonment_under_Roman_and_Medieval_settlement?email_work_card=view-paper

Van Lanen , Rowin J. and Esther Jansma, and Menne Kosian, Finding a way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An international Journal (30), 200-222. https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

Van Lanen , Rowin J. and  Bert J. Groenewoudt , Theo Spek, Esther Jansma, Route Persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands, Archaeol Anthropol Sci (2018) 10:1037–1052, DOI 10.1007/s12520-016-0431-z , https://www.researchgate.net/publication/310757823_Route_persistence_Modelling_and_quantifying_historical_route-network_stability_from_the_Roman_period_to_early-modern_times_AD_100-1600_a_case_study_from_the_Netherlands

Van Lanen, R.J., Kosian, M.C., Groenewoudt, B.J., Spek, T., Jansma, E., Best travel options: modelling Roman and early-medieval routes in The Netherlands using a multi-proxy approach. J. Archaeol. Sci. Rep. 3 (JASR), 144e159., 2015

Van Lanen, Rowin J. and Harm Jan Pierik, Calculating connectivity patterns in delta landscapes: Modelling Roman and early-medieval route networks and their stability in dynamic lowlands, Quaternary International, Volume 501, Part B, 20 January 2019, Pages 393-412, https://www.sciencedirect.com/science/article/abs/pii/S1040618216313519?via%3Dihub ; also https://www.academia.edu/34741700/Van_Lanen_and_Pierik_2017_Calculating_connectivity_patterns_in_delta_landscapes_Modelling_Roman_and_early_medieval_route_networks_and_their_stability_in_dynamic_lowlands?email_work_card=view-paper

Van Lanen, R.J., Jansma, E., Van Doesburg, J., Groenewoudt, B.J., Roman and early-medieval long-distance transport routes in north-western Europe: modelling frequent-travel zones using a dendroarchaeological approach. J. Archaeol. Sci. 73, 120e137. 2016, https://www.sciencedirect.com/science/article/abs/pii/S0305440316300978

Verhagen, P., Brughmans, T., Nuninger, L., & Bertoncello, F. , The long and winding road: Combining least cost  paths and network analysis techniques for settlement  location analysis and predictive modelling. In G. Earl, T.  Sly, A. Chrysanthi, P. Murrieta-Flores, C. Papadopoulos, I.  Romanowska, & D. Wheatley (Eds.), Archaeology in the  digital era. Papers from the 40th Annual Conference of  Computer Applications and Quantitative Methods in  Archaeology (CAA) (Southampton, March 26–29, 2013,  pp. 357–366)

Verhagen, J.W.H.P., Case Studies in Archaeological Predictive Modelling. PhD thesis. Leiden University, Archaeological Studies Leiden University (ASLU) 14, Leiden University Press, Leiden. 2007

Verhagen, J.W.H.P., Whitley, T.G., Integrating archaeological theory and predictive modeling: a live report from the scene. J. Archaeol. Method Theory 19, 49e100. 2011

Vos , Peter C,  Origin of the Dutch Coastal landscape; Long term landscape evolution to the Netherlands during the Holocene, described and visualized in national regional and local palaeographical map series, PhD Thesis Utrecht University, Utrecht, 2015,  https://dspace.library.uu.nl/handle/1874/315553

[69] Van Lanen et. al. provide basic definitions associated with this research:

Van Lanen , Rowin J. and Esther Jansma, and Menne Kosian, Finding a way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An international Journal (30), 200-222. https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

See also:

Van Lanen, R.J., Kosian, M.C., Groenewoudt, B.J., Spek, T., Jansma, E., 2016b. Route persistence. Modelling and quantifying historical route-network stability during the last two millennia: a case study from The Netherlands. Archaeol. Anthropol. Sciences 1e16. https://link.springer.com/journal/12520/onlineFirst/page/3 .

van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences. 2020;99:e14. doi:10.1017/njg.2020.12 https://doi.org/10.1007/s12520-016-0431-z

van Rowin J. and Lanen, Harm Jan Pierik, Calculating connectivity patterns in delta landscapes: Modelling Roman and early-medieval route networks and their stability in dynamic lowlands, Quaternary International, Volume 501, Part B, 2019, Pages 393-412, ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2017.03.009 .
(https://www.sciencedirect.com/science/article/pii/S1040618216313519 )

[70] See for example:

van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, and Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 30 (2015) 200–222, https://research-portal.uu.nl/ws/portalfiles/portal/258920868/Geoarchaeology_-_2015_-_Lanen__Finding_a_Way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the.pdf

van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences. 2020;99:e14. doi:10.1017/njg.2020.12 https://doi.org/10.1007/s12520-016-0431-z

van Lanen, Rowin J. and Harm Jan Pierik, Calculating connectivity patterns in delta landscapes: Modelling Roman and early-medieval route networks and their stability in dynamic lowlands, Quaternary International, Volume 501, Part B, 2019, Pages 393-412, ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2017.03.009 .
(https://www.sciencedirect.com/science/article/pii/S1040618216313519 )

Jansma, Esther and Rowin J. Van Lanen and Harm Jan Pierk, Traveling Through a River Delta: A Landscape Archaeogical Reconstruction of River Development and Long Distance Connections in the netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39, https://www.academia.edu/35074210/Jansma_E_R_J_Van_Lanen_and_H_J_Pierik_2017_Travelling_through_a_river_delta_a_landscape_archaeological_reconstruction_of_river_development_and_long_distance_connections_in_the_Netherlands_during_the_first_millennium_AD_Medieval_Settlement_Research_32_35_39

van Lanen, Rowin J. and Esther Jansma, Jan van Doesburg, Bert J. Groenewoudt, Roman and early-medieval long-distance transport routes in north-western Europe: Modelling frequent-travel zones using a dendroarchaeological approach,
Journal of Archaeological Science, Volume 73, 2016, Pages 120-137, ISSN 0305-4403,
https://doi.org/10.1016/j.jas.2016.07.010 .
(https://www.sciencedirect.com/science/article/pii/S0305440316300978 )

[71] van Lanen, R. J., Groenewoudt, B., Spek, T., & Jansma, E.. Route persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands. Archaeological and Anthropological Sciences, 10 (5), 2018, 1037–1052. https://doi.org/10.1007/s12520-016-0431-z , https://pure.rug.nl/ws/portalfiles/portal/63407562/Lanen2018_Article_RoutePersistenceModellingAndQu.pdf

R.J. van Lanen. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences, Volume 99, e14. https://doi.org/10.1017/njg.2020.12

[72] van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences. 2020;99:e14. doi:10.1017/njg.2020.12 , https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/revealing-the-past-through-modelling-reflections-on-connectivity-habitation-and-persistence-in-the-dutch-delta-during-the-1st-millennium-ad/717D79192E8EDD6B90D58B060117EB1B

Rowin J. van Lanen, Esther Jansma, Jan van Doesburg, Bert J. Groenewoudt, Roman and early-medieval long-distance transport routes in north-western Europe: Modelling frequent-travel zones using a dendroarchaeological approach, Journal of Archaeological Science, Volume 73, 2016, Pages 120-137, ISSN 0305-4403, https://doi.org/10.1016/j.jas.2016.07.010 .
(https://www.sciencedirect.com/science/article/pii/S0305440316300978 )

Jansma, Esther and Rowin J. Van Lanen and Harm Jan Pierk, Traveling Through a River Delta: A Landscape Archaeogical Reconstruction of River Development and Long Distance Connections in the netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39, https://www.academia.edu/35074210/Jansma_E_R_J_Van_Lanen_and_H_J_Pierik_2017_Travelling_through_a_river_delta_a_landscape_archaeological_reconstruction_of_river_development_and_long_distance_connections_in_the_Netherlands_during_the_first_millennium_AD_Medieval_Settlement_Research_32_35_39

van Lanen, Rowin J. and Harm Jan Pierik, Calculating connectivity patterns in delta landscapes: Modelling Roman and early-medieval route networks and their stability in dynamic lowlands, Quaternary International,
Volume 501, Part B, 2019, Pages 393-412, ISSN 1040-6182,
https://doi.org/10.1016/j.quaint.2017.03.009 .
(https://www.sciencedirect.com/science/article/pii/S1040618216313519 )

[73] van Lanen,Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, and Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, quote Page 2, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands

van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences. 2020;99:e14. doi:10.1017/njg.2020.12 https://doi.org/10.1007/s12520-016-0431-z

van Lanen, R.J., Groenewoudt, B.J., Spek, T. et al. Route persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands. Archaeol Anthropol Sci 10, 1037–1052 (2018). https://doi.org/10.1007/s12520-016-0431-z

[74] van Lanen,Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, and Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, quote Page 2, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands

[75] van Lanen, R.J., Groenewoudt, B.J., Spek, T. et al. Route persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands. Archaeol Anthropol Sci 10, 1037–1052 (2018). https://doi.org/10.1007/s12520-016-0431-z

Rowin J. van Lanen, Harm Jan Pierik, Calculating connectivity patterns in delta landscapes: Modelling Roman and early-medieval route networks and their stability in dynamic lowlands, Quaternary International, Volume 501, Part B, 2019, Pages 393-412, ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2017.03.009 .
(https://www.sciencedirect.com/science/article/pii/S1040618216313519 )

[76] van Lanen, R.J., Groenewoudt, B.J., Spek, T. et al. Route persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands. Archaeol Anthropol Sci 10, 1037–1052 (2018). Route persistence. Modelling and quantifying historical route-network stability

[77] Ai-Ping Fan, A.J. (Tom) van Loon, Ren-Chao Yang, The control of Pleistocene palaeogeography on the distribution of sandy patches in a silty Holocene lagoon (central Netherlands), Journal of Palaeogeography, Volume 11, Issue 4, 2022, Pages 565-583, ISSN 2095-3836, https://doi.org/10.1016/j.jop.2022.08.003 .
(https://www.sciencedirect.com/science/article/pii/S2095383622000852 )

Groenewoudt, Bert J., An exhausted landscape. Medieval use of moors, mires and commons in the Eastern Netherlands, In : Medieval Rural Settlement in Marginal Landscapes (pp.149-180), 2009/01/01, SN – 978-2-503-52746-8 10.1484/M.RURALIA-EB.3.1169, https://www.researchgate.net/publication/299929968_An_exhausted_landscape_Medieval_use_of_moors_mires_and_commons_in_the_Eastern_Netherlands

[78] van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences. 2020;99:e14. doi:10.1017/njg.2020.12, https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/revealing-the-past-through-modelling-reflections-on-connectivity-habitation-and-persistence-in-the-dutch-delta-during-the-1st-millennium-ad/717D79192E8EDD6B90D58B060117EB1B

van Lanen, R. J., Groenewoudt, B., Spek, T., & Jansma, Route persistence. Modelling and quantifying historical route-network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands. Archaeological and Anthropological Sciences, 10 (5), 2018, 1037–1052., https://doi.org/10.1007/s12520-016-0431-z

[79] Jansma, Esther and Rowin J. Van Lanen and Harm Jan Pierk, Traveling Through a River Delta: A Landscape Archaeogical Reconstruction of River Development and Long Distance Connections in the netherlands During the First Millennium AD, Medieval Settlement Research 32 (2017), 35–39, https://www.academia.edu/35074210/Jansma_E_R_J_Van_Lanen_and_H_J_Pierik_2017_Travelling_through_a_river_delta_a_landscape_archaeological_reconstruction_of_river_development_and_long_distance_connections_in_the_Netherlands_during_the_first_millennium_AD_Medieval_Settlement_Research_32_35_39

van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences. 2020;99:e14. doi: 10.1017/njg.2020.12 , https://www.researchgate.net/publication/347351166_Revealing_the_past_through_modelling_Reflections_on_connectivity_habitation_and_persistence_in_the_Dutch_Delta_during_the_1st_millennium_AD

Van Lanen, R.J., Groenewoudt, B.J., Spek, T. and Jansma, E. 2016b. Route persistence. Modelling and quantifying historical route network stability from the Roman period to early-modern times (AD 100–1600): a case study from the Netherlands, Archaeological and Anthropological Sciences: 1–16. https://www.researchgate.net/publication/310757823_Route_persistence_Modelling_and_quantifying_historical_route-network_stability_from_the_Roman_period_to_early-modern_times_AD_100-1600_a_case_study_from_the_Netherlands

[80] Dendrochronology, Wikipedia, This page was last edited on 19 December 2025, https://en.wikipedia.org/wiki/Dendrochronology

Jansma, E., Haneca, K. and Kosian, M. 2014b. A dendrochronological reassessment of three Roman vessels from Utrecht (the Netherlands): evidence of inland navigation between the lower- Scheldt region in Gallia Belgica and the limes of Germania inferior, Journal of Archaeological Science 50: 484–496, https://www.academia.edu/26380316/A_dendrochronological_reassessment_of_three_Roman_boats_from_Utrecht_the_Netherlands_evidence_of_inland_navigation_between_the_Scheldt_region_in_Gallia_Belgica_and_the_limes_of_Germania_inferior

Blakemore, Erin,, How are tree rings used to help date an archaeological site?, National Geographic, 8 Jul 2019, https://www.nationalgeographic.com/culture/article/how-tree-rings-date-archaeological-site?loggedin=true&rnd=1768417166750

Kristof Haneca, Katarina Čufar, Hans Beeckman, Oaks, tree-rings and wooden cultural heritage: a review of the main characteristics and applications of oak dendrochronology in Europe, Journal of Archaeological Science, Volume 36, Issue 1, 2009, Pages 1-11, ISSN 0305-4403, https://doi.org/10.1016/j.jas.2008.07.005 .
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Visser, Ronald M. “Dendrochronological provenance patterns. Network analysis of tree-ring material reveals spatial and economic relations of roman timber in the continental north-western provinces.” Journal of Computer Applications in Archaeology 4.1 2021. https://journal.caa-international.org/articles/10.5334/jcaa.79

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[81]

[82] van Lanen, Rowin J. and Menne C. Kosian, Bert J. Groenewoudt, Esther Jansma, Finding a Way: Modeling Landscape Prerequisites for Roman and Early-Medieval Routes in the Netherlands, Geoarchaeology: An International Journal 00 (2015) 1–23, https://doi.org/10.1002/GEA.21510, https://www.academia.edu/11879781/Van_Lanen_et_al_2015_Finding_a_way_Modeling_Landscape_Prerequisites_for_Roman_and_Early_Medieval_Routes_in_the_Netherlands?email_work_card=title

van Lanen RJ. Revealing the past through modelling? Reflections on connectivity, habitation and persistence in the Dutch Delta during the 1st millennium AD. Netherlands Journal of Geosciences. 2020;99:e14. doi:10.1017/njg.2020.12

The Turbulent Roman Era – The Griff(is)(es)(ith) Y-DNA Phylogenetic Gap Associated with the Meuse and Rhine River Watershed – Part Seven

This part of the story focuses on the impact of the Roman era on the absence of documented haplogroups associated with the Griff(is)(es)(ith) paternal line between the first century BCE and 300 CE in an area we now call the Netherlands and Belgium. The absence of identified YDNA subclades associated with the Griff(is)(es)(ith) YDNA paternal line in this time period may have been influenced by the effects of:

  • The changing dynamics between local Celtic and Germanic groups in the Meuse Rhine area;
  • The effects of Roman conquest and occupation;
  • Being close to or on the Roman frontier border; and
  • The late-Roman social and demographic collapse.

The Roman era did not just overlay a single Roman identity. It acted as a catalyst for a dynamic and complex set of social processes where Celtic and Germanic social groups were relocated, subjugated, blended or destroyed. This lead to the formation of new group identities that would define the post-Roman, early-medieval landscape of the region, ultimately contributing to the development of the Frankish kingdom. 

Rural communities and social groups underwent major transformations in this period of time. The violently disruptive Caesarian Roman conquest of the area in the first century BCE led to substantial demographic losses and trauma for indigenous populations. The subsequent large-scale migration and managed resettlement of Germanic groups resulted in new tribal configurations and ethnogenesis (the process by which a new ethnic identity emerges from existing groups, often through migration, interaction, and cultural blending). The integration of rural populations into the Roman military system, with extensive recruitment for auxiliary forces, also led to the accelerated diffusion of Roman military culture and altered local social structures. [1]

The Changing Dynamics Between Indigenous Groups in the Meuse Rhine Watershed Area Pre-Roman Era

It is highly likely that the changes in Celtic and Germanic group composition and the impact of the Roman incursion in the Meuse Rhine watershed area during this time period had a major impact on the continued absence of documented haplogroups in the YDNA Griff(is)(es)(ith) YDNA phylogenetic line. It probably contributed to the resultant and reinforced pattern of a phylogenetic tree with long branches between identified haplogroups with minimal subclade formation. [2]

High levels of intergroup competition, associated with migration or conflict, led to the physical displacement or decline of male lineages. This resulted in observable changes in haplogroup prevalence over time in the Meuse Rhine watershed region. Social and culturally driven group processes (i.e.the Roman incusion and subjagation) resulted in specific Y-DNA lineages (clades) dying out more frequently than would be expected by random chance.

This altered the overall structure of the YDNA phylogenetic tree, potentially leading to the loss of entire branches that, in the absence of such social pressures, might have persisted. Interaction and conflict among social groups acted as a potent filter on Y-DNA transmission, making Y-DNA phylogenetic trees a reflection not only of biological mutation and migration but also of the cultural and social history of the populations they represented.

Prior to the Roman impact (pre-first century BCE), the area between the Meuse and Rhine rivers was inhabited by a mix of Celtic and Germanic groups, some displaying transitional identities and social pattterns due to centuries of interaction and migration. The identity and culture in the Rhine and Germania regions during this period were highly fluid, marked by ongoing processes of migration, assimilation, and hybridization among small groups rather than fixed ethnic divides and geographical boundaries.

A comparative review of maps depicting Germanic and Celtic groups during this time peiod show a wide range of variable depictions of indigenous groups. These maps often differ significantly in their representation of tribal boundaries, ethnic affiliations, and even the names assigned to specific groups. See illustrations one through six as examples.

Illustration One – Six: Examples of Variability of Identified Local Indigenous Groups in the Meuse Rhine Watershed Area and Outlying Areas

Illustration One: Tribal map of the Lower Rhine frontier zone in the Caesarian Era

Click for Larger View | Source: Roymans N, Derks T, Heeren S (2020) Roman Imperialism and the transformation of rural society in a frontier province: diversifying the narrative. Britannia 51:265–294. https://www.cambridge.org/core/journals/britannia/article/roman-imperialism-and-the-transformation-of-rural-society-in-a-frontier-province-diversifying-the-narrative/BF7FA75997E14A1B7BA4241E5DC981C3

Illustration Two: Map of Germanic and Celtic Tribes circa 52 BCE

Click for Larger View | Source: Modification of map by P L Kessler, Map of Celtic and Germanic Tribes,05 March 2015, World History Encyclopedia,https://www.worldhistory.org/image/3687/map-of-celtic-and-germanic-tribes/

Illustration Three: The Germanic Tribal World of the Provincial Roman Period between 50 BCE to 300 CE

Click for Larger View | Source: A section of a map created by Arch.-Stud. A.P., The Germanic Tribal World of the Provincial Roman Period between 50 BC to 300 AD, 7 Apr 2023, Wikimedia Commons, https://commons.wikimedia.org/wiki/File:Germanic_Tribes_in_the_Roman_Imperial_Period.png

Illustration Four: Map of the Germanic tribes, circa 15 BCE

Click for Larger View | Source: Jason Abdale, Map of the Germanic tribes, circa 15 BC, 26 Jul 2013, Dinosaurs and Barbarians,https://dinosaursandbarbarians.com/2013/07/26/map-of-the-germanic-tribes-circa-15-bc/

Illustration Five: Roman provinces & tribes in vicinity of Gallia c 58 BC, prior to Gallic Wars

Click for Parger View | Source: Modified version of a map by Feitscherg, Map of Roman provinces & tribes in vicinity of Gallia c 58 BC, prior to Gallic Wars,2 Sep 2019, Wikimedia Commons,https://commons.wikimedia.org/wiki/File:Map_Gallia_Tribes_Towns-la.svg

Illustration Six: Map of Celtic, Germanic and Iberian tribes before Roman expansion

Click for Larger View | Source: Modified version of map documented in a 2017 reddit post by ironandrewoods, I drew a map of the pre-Roman peoples of Western Europe and North Africa! [3170 × 5042] [OC], https://www.reddit.com/r/MapPorn/comments/6u61sl/i_drew_a_map_of_the_preroman_peoples_of_western/

There are a number of reasons for this variability. Many maps are based on fragmentary and sometimes contradictory ancient sources, such as Roman and Greek accounts, which themselves were often incomplete or biased.​ The boundaries and identities of tribes were fluid, with frequent migrations, alliances, and assimilations, making it difficult to draw fixed lines on a map.

Modern mapmakers often extrapolate from limited evidence, leading to different interpretations and sometimes speculative placements of tribes. Modern maps often rely on a combination of archaeological evidence, linguistic data, and historical texts, but these sources do not always align, resulting in differing interpretations of tribal territories and ethnic identities.

Scholars debate the extent to which ethnic or tribal identities were fixed or fluid, with some arguing for a more dynamic and overlapping picture rather than clear-cut divisions. The lack of consensus on the precise locations and affiliations of many tribes means that different maps may show significant discrepancies, especially in regions like the Rhine-Meuse watershed where Germanic and Celtic groups interacted closely.

Illustration seven (below) reflects the general location of the main Celtic and Germanic groups in the Meuse Rhine watershed area prior to Roman occupation. The map is the result of research conducted by Fanny Martin. Her research was aimed at establishing
a reliable geographic and chronological framework based on material culture and territorial dynamics of the late Iron Age tribes in the middle Meuse basin on the eve of the Roman Conquest. [3]

Illustration Seven: Tribal Groups in the Meuse Rhine Watershed Area Prior to Roman Occupation

Click for Larger View | Modification of original image found in: Martin, F. (2019). Recent research on material culture and territorial dynamics of late Iron Age tribes in the middle Meuse basin, on the eve of the Roman Conquest, Abb. 10 Proposed localization of the main Celtic tribes of the area in : Spätlatènezeitliche und frühkaiserzeitliche Archäologie zwischen Maas und Rhein, Bonn, 323-334. https://www.academia.edu/43850047/Martin_F_2019_Recent_research_on_material_culture_and_territorial_dynamics_of_late_Iron_Age_tribes_in_the_middle_Meuse_basin_on_the_eve_of_the_Roman_Conquest_In_Spätlatènezeitliche_und_frühkaiserzeitliche_Archäologie_zwischen_Maas_und_Rhein_Bonn_323_334

Martin’s methodology for mapping late Iron Age tribes in the middle Meuse basin is noteworthy. It is centered on a detailed, comparative analysis of the distribution of material culture (primarily ceramics, glass bracelets, salt containers, fibulae, and coinages) and settlement patterns (the modalities and distribution of housing, farming evidence, defensive settlements, funeral practices or cave occupation) within a newly established chronological framework. The archaeological data was compared with historical sources to observe the integration of local populations into the Roman sphere of influence. I added the approximate location of Germanic groups east of the Rhine to the map.

Based on the approximate migratory path of the Griff(is)(es)(ith) paternal genetic line, it is possible a number of generations of the Griff(is)(es)(ith) line were associated the Treveri Celtic group. Depending on the rate of migratory movement over time, generations of the genetic line may also have been associated with the Eburones.

Between 300 BCE and 1 BCE, the area between the Meuse and Rhine Rivers underwent significant changes, with a changing mix of Celtic and Germanic groups inhabiting the area, including some with hybrid origins or shifting allegiances due to group migrations.

Migration and assimilation in the Meuse-Rhine region before the Roman era led to dynamic changes in local tribal groups, with tribal identities shifting, merging, and evolving in response to population movement and intergroup contact. Tribal group migration resulted in tribal splits, hybridization and changes in group characteristics. Migration and assimilation transformed local tribal communities and tribal identity in the Meuse-Rhine region from static designations to fluid, ‘negotiated identities’. [4]

Internal disputes, environmental pressures, or external threats spurred subgroup migrations—like the Batavians splitting off from the Chatti and settling in the Rhine delta. Another example is the Ubii, allied with the Romans, moving from the east bank of the Rhine to the west bank, filling a geographical vacuum where the annihilated Eburones lived. These relocated groups often adopted new names and distinct identities tailored to their new socio-political environment, sometimes preserving older tribal names only superficially while forming completely new communities.[5]

Migrants and resident populations engaged in broad facets of acculturation such as intermarriage, shared burial rites, and exchange of technologies which led to changes in tribal identification. Dress styles and funerary customs, for example, became markers of new group identities, not indicators of longstanding ethnic continuity.[6]

‘Ethnic’ characteristics, such as language, material culture, or place-names were changed with local groups and newcomers often combining or abandoning traditions for social advantage. This ‘adaptive negotiation’ generated new identities with names linked to older tribal entities but whose membership and meaning were entirely transformed. [7]

The process of tribal assimilation was not unidirectional. Groups negotiated and adapted cultural elements for prestige and belonging. Immigrant “barbarians” and original inhabitants adopted or performed traditions to establish social distinction, producing a blended, hybridized population and new ethnic groupings. [8]

As new communities formed, they often retained ancient tribal names—such as Franks, Batavians, or Chatti—but these now referred to socially and genetically mixed groups divergent from their predecessors. [9] Recent genetic and archaeological studies reveal that these processes led to mosaic populations—emphasizing local continuity alongside significant input from newcomers, further destabilizing clear-cut tribal labels. [10]

Archaeological and linguistic evidence shows considerable hybridization and cultural mixing. Many tribal names and material cultures east or west of the Rhine (e.g., Chatti, Batavi, Eburones, Nemetes) displayed La Tène (Celtic) signatures even after initial “Germanic” migrations, leading scholars to recognize transitional or hybrid populations with changing language, traditions, and identities. [11]

Repeated waves of migration, assimilation, and sociopolitical restructuring led to evolving ethnic boundaries. For example, tribes that began as Celtic-speaking could be subsumed by groups adopting Germanic languages and material cultures through intermarriage or power shifts. This is reflected in archaeological records and analyses of personal and place-names, all of which document the fuzzy boundaries between groups. [12]

The Identification of Indigienous Groups in the Pre-Roman and Roman Era

Recent research in the last fifty years on indigenous communities in the Meuse-Rhine watershed has critically reconsidered the use of the term “tribe.” The concept is increasingly viewed as problematic because it reflects nineth-century sociopolitical models more than the actual archaeological or ancient realities. [13]

The naming of Celtic and Germanic tribes in the Meuse-Rhine area before and during the Roman era has been fraught with difficulty due to both scholarly and historical ambiguities about ethnic identities, linguistic affiliations, and shifting cultural frontiers.

The term “tribe” was largely imposed by Roman and later classical authors, as well as modern ethnographers, onto diverse and dynamic local groups whose social, political, and cultural organization do not match the simplified, static model implied by “tribe.” Archaeologists, anthropologists, and historians now recognize that local identities were more fluid, overlapping, and constructed in response to both local and imperial pressures, rather than being fixed or primordial entities. [14]

An attempt to describe the environmental (and social) aspects of the prehistory and early history of the lowland rivers Rhine and Meuse is a confrontation with a poignant lack of information. How little do we know from that period? The most reliable records come from geological, palaeogeographical and palynological research, and from scanty archaeological excavations. The first written sources date back to the Roman period, but these are subjective, often second-hand observations of educated military men and historians. After the Roman period, until roughly ad 800–900, again, there is a great lack of written documentation.[15]

Beginning around the late second to early first century BCE, the Meuse–Rhine region became home to several Celtic or partly Celtic tribes belonging to the Belgic cultural sphere. These tribes occupied territories that bridged Celtic and early Germanic zones, [16] forming a cultural frontier that Julius Caesar later described in his Commentarii de Bello Gallico. [17]

Read more on Ethnicity, Tribes and Archaeology

Ethnicity, Tribes and Archaeology

In many ways, ethnic identity has remained the elephant in the room in many aDNA-driven narratives. While there have been efforts to revise nomenclature to better reflect criticisms of the culture concept and its inherent assumptions about collective identity, in practice many models still work on the assumption that DNA signatures and experienced identities as reflected in material culture should largely coincide.” [18]

Scholars consistently caution against equating specific artifacts or burial customs with fixed ethnic identities, emphasizing the dynamic, negotiated nature of ethnicity in the archaeological record.

Ethnicity is increasingly studied as an active, socially constructed phenomenon, not merely a biological or cultural inheritance, resonating with trends in both historical and prehistoric research.

Studies of ethnicity in complex societies highlight both the difficulties and insights that archaeological approaches contribute, advocating a critical stance on linking material culture directly to ethnic groups.

The concept of ‘tribal society’ is one of the most prominent and popular ‘anthropological’ notions of our time, yet within western social and cultural anthropology it has been largely abandoned as a sociological category. . . . In the nineteenth century, the term tribe was woven into the theories of primitive society governed by the principles of ‘kinship’ proposed by the emerging social sciences. . . . By the beginning of this century ‘the tribe’ had been widely discredited as an analytical term outside some specialized fields such as theories of early state formation. It is now commonly considered an ethnographic, rather than an analytical, term. [19]

Illustration Eight: Germanic Forces Crossing the Rhine 406 CE

Germanic Forces Cross the Rhine, 406 CE
Click for Larger View | Source: modified version of an original image from Ancient Warfare Magazine/ Karwansaray Publishers. It was uploaded by Arienne King and published on 13 September 2020 at the World History Encyclopedia website. “Germanic Forces Cross the Rhine, 406 CE, World History Encyclopedia, https://www.worldhistory.org/image/12731/germanic-forces-cross-the-rhine-406-ce/

The Rhine River served as a de facto cultural boundary between Celtic Gaul and Germania. Table one provides a list of the key local groups, or tribes as defined by the Romans, in the Meuse Rhine watershed area during the late second to early first century BCE . [20]

Table One: Celtic-Germanic Groups West of the Rhine River First Centruy BCE

Roman NameDescription
EburonesThis was a large tribe situated between the Rhine and Meuse rivers, in the area of modern-day Limburg (southern Netherlands), eastern Belgium, and the German Rhineland. Caesar claimed to have virtually exterminated them after their revolt in 53 BCE, though many likely lived on and were later known as the Tungri.
Aduatuci/
Atuatuci
This grouup lived in the area of modern-day Tongeren, Belgium, between the Scheldt and Meuse rivers.
NerviiKnown for their fierce resistance to Roman rule, the Nervii were a powerful Belgic ‘tribe’ located north of the Eburones.
SegriThe Segni are generally presumed to have dwelled in the Luxembourgish and Belgian Ardennes. Their territory was located between that of the Treveri and the Eburones, indicating that they settled not far from the Condrusi,
TreveriSituated in the left bank of the Middle Rhine region, around the Ardennes and Eifel areas (modern Luxembourg and the surrounding regions), they had a mixed Celtic-Germanic culture.
Condrusi/ CondrusThis was a small, left-bank Celtic-Germanic group in the Middle Rhine region. Like the Segni, their territory was located between that of the Treveri and Eburones. At the time of Caesar’s conquest of the region in the mid-first century BCE, they lived as clients of the Treveri.
Caeroesi/ CaerosiThe Caeroesi were a left-bank, Celto-Germanic tribe located in the Eifel-Ardennes area.
MenapiiThe Menapii were a Celtic tribe with Germanic influence, located primarily south of the Meuse river. 

What Celtic-Germanic tribe are We from?

As indicated in the first part of this multipart story, the estimated migratory path between haplogroup G-FGC716 and haplogroup G-Z6748, the two endpoints associated with this phylogenetic gap, was visually structured through the use of the FamilyTreeDNA Globe Trekker program. In FamilyTreeDNA’s GlobeTrekkeprogram, corridor paths represent  probabilistic zones indicating the most likely routes Y-DNA lineages followed between ancestral haplogroup locations. 

Illustration nine depicts the migratory corridor where the Griff(is)(es)(th) lineage most likely traveled, based on 95 percent probability, through time rather than a single deterministic line. It also indicates the general area where various Celtic and Germanic groups resdes during the pre-Roman era.

Illustration Nine: Tribal Map of the Lower Rhine frontier Zone in the Caesarian Period circa 50 BCE

Click for Larger View | Source:Roymans N, Derks T, Heeren S (2020) Roman Imperialism and the transformation of rural society in a frontier province: diversifying the narrative. Britannia 51:265–294. https://www.cambridge.org/core/journals/britannia/article/roman-imperialism-and-the-transformation-of-rural-society-in-a-frontier-province-diversifying-the-narrative/BF7FA75997E14A1B7BA4241E5DC981C3

Since the rate of northwest movement of the undocumented lineages associated with the Griff(is)(es)(ith) YDNA migratory path are not known, the ability to pinpoint the location of roughly ninety generations of ancestors living along this migratory zone is uncertain. Based on the date ranges associated with the two endpoints of the phylogenetic gap (see illustration ten below), as previously stated, it is likely that the generations that lived during the pre-Roman era may have lived in an area where either the Eburones or the Treveri lived.

It is perhaps more likely the generations that lived in this time period were in the territory that was occupied by the Eburones.

Illustration Ten: Estimated Birth Date Ranges of the Ancestors Associated with the Haplogroup Endpoints of the Phylogenetic Gap Based on a 95 Percent Probability Rate

Click for Larger View | Source: FamilyTreeDNA, Scientific Details on Haplogroups G-FGC7516 and G-Z6748

The Eburones lived in an area broadly situated between the Ardennes and Eifel region in the south, and the Rhine-Meuse delta in the north. Their territory lay east of the Atuatuci (themselves east of the Nervii), south of the Menapii, and north of the Segni and Condrusi (themselves north of the Treveri). To the east, the Sugambri and Ubii were their neighbours on the opposite bank of the Rhine. When the Germanic Tencteri and Usipetes crossed the Rhine from Germania in 55 BC, they first fell on the Menapii and advanced into the territories of the Eburones and Condrusi, who were both “under the protection of ” the Treveri to the south.[21]

The Eburones were a Belgic-Germanic tribe who lived in the northeastern part of Gaul during the Roman conquest, in what is now southern Netherlands, eastern Belgium, and the German Rhineland, situated between the Rhine and the Meuse rivers.

Caesar explains that the heartland of the Eburones was between the Meuse and Rhine, which probably is more or less identical to the Belgian and Dutch provinces called Limburg, and the western part of Nordrhein-Westphalen. In any case, it was north of the Ardennes. South of these old mountains lived the Treverans, of whom the Eburones were a client-tribe, which was protected by the mightier tribe.[22]

The Eburones are primarily known for their rebellion against Julius Caesar during the Gallic Wars (54–53 BCE), which led to the near-extermination of their people by the Roman army. Their revolt, led by Ambiorix, resulted in the destruction of 15 Roman cohorts at Aduatuca before the Romans retaliated ( an artistic rendtion of this battle is portrayed in illustration twelve below). Following the revolt, Caesar led a punitive campaign against the Eburones, resulting in their near-complete annihilation. The tribe effectively disappeared from historical records afterward, with new tribes eventually settling their former territory. [23] Whether any significant part of the population lived on in the area as Tungri, the tribal name found here later, is uncertain but considered likely. [24]

A significant portion of the Eburones population appears to have been devastated by Julius Caesar’s retaliatory campaigns after their rebellion, as archaeological evidence demonstrates a dramatic demographic decline and settlement abandonment in the Eburonean core area between the Meuse and Rhine rivers in the mid-first century BCE. However, archaeological and historical studies indicate that although many settlements were depopulated or destroyed, the territory was not completely emptied of inhabitants, and some degree of population continuity is supported by later reoccupation evidence. [25]

Illustration Eleven: Model of the Eburonian settlement at Hambach-Niederzier, which was abandoned after the mid-first century BCE

Click for Larger View | Source: Jona Lendering, Model of the Eburonian settlement at Hambach-Niederzier, Photo source – Marco Prins, Rheinisches Landesmuseum, Bonn , Livius.org, https://www.livius.org/pictures/germany/hambach-niederzier/hambach-niederzier-eburonian-village-model/

There is archaeological evidence for their survival and discontinuity as a group in the area. Archaeological surveys show a sharp break or reduction in habitation following the rebellion, but traces of subsequent reoccupation or survival, particularly in the western parts of Eburonean territory, are evident later in the Roman period.​ Some fortified sites, possibly used as refuges, lack evidence of use after the mid-first century BCE, indicating localized extinction. However, refuge was likely sought in inaccessible regions like the Ardennes, swamps, or coastal islands.​ The rapid return of settlement in parts of the region during the later first century BCE and early first century CE suggests outsiders (likely Germanic settlers invited by the Romans) filled some of the demographic vacuum left by the devastation. [26]

Illustration Twelve: Artist’s impression of the battle between the Eburonean king Ambiorix and the Roman army in 54 BC

Click for Larger View | Source: Roymans, N. and Fernández-Götz, M., (2015): Fire and Sword. The archaeology of Caesar’s Gallic War. Military History Monthly 56: 52-56. https://www.academia.edu/12866878/Fire_and_Sword_The_archaeology_of_Caesar_s_Gallic_War

Caesar’s claim that the Eburones and their name were eradicated is likely exaggerated. Ancient historians (notably Tacitus) describe the emergence of the Tungri, who settled in the same region, as possible successors or absorbers of the remaining Eburonean elements.[27]

Modern scholarship, such as that of Nico Roymans, Ton Derks, Stijn Heeren and others suggest that the disappearance of the Eburones from historical records may reflect Roman policies of land confiscation and deliberate erasure of ethnic labels, rather than true population extermination.​ The formation of new tribal entities (like the Tungri) likely involved the assimilation of surviving Eburonean groups, people from neighboring tribes, and new migrants, supported by some archaeological continuity in the landscape. Landscape archaeology, rather than genetics, currently provides the most persuasive evidence for both a severe demographic crisis and eventual population mixing and cultural transformation in the area after the Gallic Wars. [28]

Impact on Local Groups After the Gallic Wars

Archaeological surveys in the Meuse–Rhine region show abrupt discontinuity in settlement patterns during the first century BCE, specifically around the time of Caesar’s campaigns. Stratified archaeological deposits, weapon caches, and mass graves (see illustration thirteen) have been linked to documented events such as Caesar’s slaughter of the Usipetes and Tencteri, with forensic examinations of human remains (perimortem trauma, carbon dating) supporting the association (see illustration thirteen). The landscape was partly depopulated due to massacre and displacement, and occupation patterns only resumed under Romanization and later Germanic migrations. [29]

Illustration Thirteen: Human remains from a battle-related find complex dredged from the River Meuse at Kessel-Lith (the Netherlands), probably linked to Caesar’s massacre of the Germanic Tencteri and Usipetes in 55 BC

Click for Larger View | Source: Manuel Fernández-Götz, Dominik Maschek and Nico Roymans, The Dark Side of the Empire: Roman Expansionism between Object Agency and Predatory Regime, https://www.researchgate.net/publication/346718218_The_dark_side_of_the_Empire_Roman_expansionism_between_object_agency_and_predatory_regime

An article by Roymans, Derks, and Heeren challenges conventional narratives of Roman ‘romanisation’ of northwestern europe (see sidebar discussion on Romanization versus the Imperial Agent model of ethnogenesis) by focusing on the often overlooked exploitative and repressive aspects of Roman rule, especially in rural frontier societies. The work highlights the crucial role that Roman imperial power played in shaping—and sometimes destabilizing—indigenous groups and rural communities and their social dynamics. [30]

The writers argue that Roman imperial agency in the emerging Roman province of Germania Inferior was direct, dynamic, and often disruptive, fundamentally reshaping rural society through mechanisms that went far beyond ‘romanisation’ as peaceful cultural diffusion. Imperial policies were implemented as top-down interventions, orchestrating violence, population movements, and administrative restructuring to serve the needs of the Roman Empire.

Roman authorities actively engineered mass violence, depopulation, and forced migration during the initial conquest phase, leaving the region traumatized and its population dramatically reduced. (See illustration thirteen above and fourteen below).

The post-conquest era saw imperial actors organize the large-scale settlement of new Germanic groups, using land allocation and tribal reorganization as a means of exercising control and maintaining stability in frontier zones.

The Romanization Model Versus the Imperial Agent Model of Ethnogenesis

Roymans et al.’s article diverges sharply from the traditional archaelogical Romanization model, providing a critique rooted in imperial agency and highlighting exploitation, violence, and engineered social transformation, especially in Germania Inferior.

The traditional Romanization paradigm interprets provincial change as largely a process of cultural assimilation and acculturation, where local elites emulate Roman ways and gradually spread these practices to the broader population, with a strong focus on urbanism, elite culture, and the ‘civilizing’ effects of Rome.

Key Points of Contrast:

  • Traditional models highlight voluntary adoption and negotiation, viewing rural transformation as peaceful and evolutionary.
  • Roymans et al. argue rural societies were fundamentally transformed by direct imperial actions, involving large-scale violence, population displacement, and elite reshuffling, with Romanization often occurring as a byproduct of imperial policies rather than as a result of indigenous choice or gradual acculturation.
  • The paper positions Germania Inferior not as a passive recipient of civilization, but as a ‘laboratory for imperial social engineering’ and the exercise of Roman power.

Roymans et al. insist that understanding frontier rural change requires revealing Roman expansion’s “dark side” and dismantling the simplistic, optimistic narrative of Romanization as peaceful assimilation.

Read More on Roman frontier archaeology and studies on imperial power and rural transformation

The violence of the Gallic Wars led to the near-genocide of certain local groups, dramatically altering social organization and local ethnic identities. Literary and archaeological evidence suggest an almost complete purge of prior inhabitants in some areas, but not total extinction—remnants persisted, with subsequent resettlement by new peoples during the Roman and Early Empire periods. (See illustration fourteen) [31]

Illustration Fourteen: Distribution of cases of genocide in Gaul described by Caesar in his Commentarii

Click for Larger View | Source: Modified veersion on map,Distribution of cases of genocide in Gaul described by Caesar in his Commentarii, in Roymans, N. 2019: ‘Conquest, mass violence and ethnic stereotyping: investigating Caesar’s actions in the Germanic frontier zone’, Journal of Roman Archaeology 32, 443, see note 26c

Roymans and his colleagues use a series of interrelated case studies focused on the Lower Rhine frontier—specifically the region of Germania Inferior—to illustrate the transformative effects of Roman imperial expansion, population movement, and rural reordering.

(1) Mass Depopulation and Repopulation of Germania Inferior: The researchers analyze the near-total depopulation of the countryside in the southern Netherlands and northern Belgium (Germania Inferior) following Caesar’s conquest, using archaeological settlement discontinuities and written sources about forced deportations as evidence.

One argument for taking (Caesar’s written) account of his destructive activities seriously . . . is the radically-altered tribal map of the Lower Germanic frontier in the early post-conquest period, combined with reports of the substantial settlement of immigrant groups from the E(ast) bank of the Rhine (illustration fifteen below). This settlement of new groups — the Batavi are even said to have moved to uninhabited land (Tac., Hist. 4.12: vacua cultoribus) in the Dutch river area — implies a phase of demographic decline in the preceding period.[32]

Illustration Fifteen: Tribal map of the Lower Rhine frontier zone in the early Imperial period (c. 27 BCE – 193 CE)

Click for Larger View | Source: Roymans N, Derks T, Heeren S (2020) Roman Imperialism and the transformation of rural society in a frontier province: diversifying the narrative. Britannia 51:265–294. https://www.cambridge.org/core/journals/britannia/article/roman-imperialism-and-the-transformation-of-rural-society-in-a-frontier-province-diversifying-the-narrative/BF7FA75997E14A1B7BA4241E5DC981C3

Most tribes in the Germanic frontier zone were confronted with another variant of mass violence: the scorched-earth policy. This policy was practised against groups who avoided an open battle and turned to guerrilla warfare, such as the Menapii, Morini, Sugambri, and above all the Eburones in the years after their revolt in 54 B.C. The Roman strategy was the large-scale (i.e., in actions involving several legions) burning to the ground of settlements, taking of prisoners, carrying off cattle, and destroying the harvest. Because of their sheer scale and systematic planning, the scorched-earth campaigns will have had a disastrous short-term impact on the subsistence and demography of the indigenous populations.[33]

(2) Augustan Reordering and Migration: The authors present the influx of new Germanic groups during the Augustan era as a case of strategic repopulation. This includes the administrative creation of new tribes and the integration of war bands and their leaders into existing civitates, with many of these groups recruited as auxiliaries for the Roman military.

In the Roman era, a civitas was an autonomous local administrative unit, essentially the capital town of a pre-Roman tribe or a large ‘tribal’ area. They were crucial in frontier regions because they served as centers for Roman tax collection, economic activity, and cultural administration, extending Roman influence through local elites rather than direct rule. Each civitas had a town council (curia or ordo) and was responsible for governing the surrounding rural territory. [34]

“(T)he post-conquest period, and in particular the age of Augustus, can be described as a formative phase for the Lower Germanic frontier zone, characterised by the influx of new Germanic groups from the eastern bank of the Rhine, the formation of new tribes and the first administrative ordering of the military district of Germania inferior. This resulted over the course of the first century in a fundamental reorganisation of the tribal map.[35]

Based on the written sources, the post-conquest period, and in particular the age of Augustus, can be described as a formative phase for the Lower Germanic frontier zone, characterised by the influx of new Germanic groups from the eastern bank of the Rhine, the formation of new tribes and the first administrative ordering of the military district of Germania inferior. This resulted over the course of the first century in a fundamental reorganisation of the tribal map compared to that of the Caesarian period . . . . Germania inferior most likely had six civitates . . .  : the Ubii around Cologne, the Cugerni near Xanten, the Tungri with Tongres as their centre, the Batavians around Nijmegen, the Cananefates with their capital at Voorburg and the Frisiavones in the coastal area of Zeeland. In addition, we hear about a number of smaller tribes, such as the Texuandri, Baetasii and Sunuci, which were probably attributed (as pagi?) to one of the civitates mentioned.” [36]

Illustration Sixteen: Recruitment of Indigenous Population in Roman Military for Roman Auxiliary Units

Click for Larger View | Source: FIG. 6. Overview of pre-Flavian ethnic recruitment by Rome in Germania inferior and Gallia Belgica (data after Alföldy Reference Alföldy1968): (A) civitates used for the conscription of auxiliary units; (B) ala; (C) cohors, inRoymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294

(3) Ethnic Reconfiguration and Political Constructs: Roymans, Derks and Heeren draw attention to the use the phenomenon of ‘created tribes’, such as the Batavians, as case studies for political (rather than purely ethnic) identity formation driven by Roman needs for human resources and political control.

The Ubii were moved from the right to the left bank of the Rhine (the Cologne region) by Agrippa in 39/38 or 20/19 B.C. The Batavi – a subgroup of the Chatti who lived in modern Hesse – were allowed to settle in an area said to be empty (vacua cultoribus) in the eastern half of the Dutch Rhine delta at some time between Caesar’s departure from Gaul and the arrival of Drusus in about 15 B.C. In 8 B.C. Tiberius transferred a group of 40,000 subject Sugambri and Suebi to the Gallic bank of the Rhine, where they were probably settled in sparsely populated regions; this is the only case for which we have information about the size of an immigrant population. The Cugerni and the Baetasii in the Xanten territory are often regarded as potential descendants of this immigrant group. The Cananefates in the Dutch coastal area had a close relationship with the Chauci, and probably originated in the North Sea coastal area, and the Frisiavones are often considered the descendants of a group of Frisians transferred by the Roman general Corbulo in 47. Although this is not always explicitly mentioned, it is clear that the land allocations to Germanic groups were often directed or at least sanctioned by the Roman authorities, thereby continuing a long tradition of rearranging both land and people in newly conquered areas. [37]

Roman conquest replaced Celtic political structures with Roman administrative systems, and Roman colonies were established to consolidate control. Land grants to Roman citizens in Celtic territory spread Roman culture and population, accelerating Romanization. The term “romanization” or cultural Rominzation refers to a historical process, such as the adoption of Roman culture, law, and language by conquered peoples in the Roman Empire. This involved cultural assimilation and integration into the empire’s systems, often for political or economic advantage.

From the days of Augustus until the 5th century AD the basic units of the Roman imperial administration were the civitates, consisting of an urban center and its surrounding countryside. The landowning elite controlled the administration of the cities as well as the rural peasantry. [38]

Despite Roman conquest and dominance, Celtic culture did not entirely vanish in the Roman Empire. Celts selectively adopted Roman customs, laws, and governance while retaining aspects of their identity. During the Roman era and Post Roman era, Celtic culture became mainly restricted to peripheral areas like Ireland, western and northern Britain, and Brittany. [39]

Given the general migratory path of the descendants of the YDNA genetic line of the Griff(is)(es)(ith) family, it is likely that generations leading up to the endpoint of this phylogenetic gap were associated with Germanic indigenous communities.

Table Two: Summary of Roman Impact on Celtic and Germanic Indigenous Groups

AspectGermanic TribesCeltic Tribes
Roman InteractionMilitary threat, conflict; then relocation and utilized for human resources and political control Conquest, depopulation, territorial loss, absorbtion into Germanic groups, Romanization
Settlement TypeRoman military colonies and trading settlements near frontiersRoman colonies and military outposts in conquered territories
Territorial ImpactFixed borders replacing fluid tribal territories; westward migration into former Roman landsTraditional tribal lands fragmented or absorbed into Roman provinces
Political ChangesLarger confederations, Roman-style governanceReplacement of tribal structures by Roman government
Cultural InfluenceAdoption of Roman law, ChristianitySelective adoption of Roman customs
Migration & SettlementPushed westward by Huns, formed successor statesSpread of Roman colonies, confinement to fringes of Roman Empire
Long-Term OutcomeFusion of Roman and Germanic elites creating medieval European nobilityRetention in remote regions of Empire; cultural survival with Roman elements

The Enduring Impact of the Roman Limes on Indigenous Groups along the Rhine River in Muese-Rhine Watershed Area

The Roman Empire transformed the Meuse–Rhine river economy from loosely connected agrarian communities into a frontier zone tied into imperial supply, trade, and monetary systems, but this integration was uneven over time and collapsed again from the late third century CE onward. Military provisioning, recruitment, and riverine transport were the core drivers of change, with Batavian and other local communities becoming heavily dependent on the army market while retaining a localized rural economy. [40]

Archaeological evidence from settlement patterns, route networks, and landscape use indicates that the arrival of Roman rule led to major enduring demographic and economic shifts in the region. New forts, river ports, villa landscapes, and Romanized settlements appeared along the Rhine and Meuse Rivers and the region, reflecting both administrative restructuring and increased integration of the land and people into Roman economic systems. [41]

The Roman era in northwestern Europe primarily lasted from the late first century BCE to the fifth century CE, with a sustained presence developing in areas like Gaul and Britannia. Roman conquest and control varied, with provinces established along the Rhine River by the late first century CE and Roman withdrawal from Britain around 410 CE. While Roman influence was present earlier, a sustained military and administrative presence developed after 27 BCE. Roman forces under Augustus waged campaigns into Germanic territories. After the Battle of the Teutoburg Forest in 9 CE, they retreated and consolidated control west of the Rhine River. [42]

Illustration Seventeen: The Lower Limes Roman Fronter

Click for Larger View | Source: Fig. 1. The Lower German Limes: World Heritage Sites (D/NL) in Gerlach R, Meurers-Balke J, and Kalis AJ. The Lower Rhine (Germany) in Late Antiquity: a time of dissolving structures. Netherlands Journal of Geosciences, Volume 101, e14. https://doi.org/10.1017/njg.2022.11

The Rhine served as a frontier, cultural and political boundary. Roman engineering along the Rhine river and the development of roadways during the first and third centuries CE influenced the environmental configuration of the region.

The Roman limes along the northern border of the Meuse-Rhine watershed acted as both ‘a barrier and a bridge’, reshaping indigenous society through ongoing cycles of integration, resistance, and transformation, as documented by archaeological and anthropological studies. [43]

The Limes refers to the fortified boundaries or frontiers of the Roman Empire that marked the limits of Roman territory and played a vital defensive, administrative, and cultural role throughout antiquity (see illustration seventeen). Originally a Latin term for a path or boundary line, it evolved to describe elaborate systems of border defenses consisting of walls, ditches, ramparts, palisades, watchtowers, forts, and military roads positioned along the Empire’s edges. Forts and towns were built on undercut river meanders with stabilized banks (e.g., Packwerk structures at Xanten) to ensure year-round harborage. [44] 

As reflected in illustration seventeen above, the approximate migratory path of the Griff(is)(es)(ith) paternal YDNA genetic line generally mirrors the contour of the Roman frontier Limes boundary. The Limes transitioned from a defensive line to a trade corridor by the third century CE, guarded via mobile patrols as water based activity increased.

Along the Roman frontier, especially near the Rhine and Danube rivers, Roman military colonies and fortifications were established to secure borders. These settlements marked a shift from tribal, fluid territories to more controlled boundary influenced by Roman presence.

Living along the Roman limes was envisioned—according to archaeological and anthropological studies—as an experience of liminality, shaped by borderland dynamics, military presence, economic integration, and a mix of cultural groups. [45]

Roman trade and cultural exchange led some Germanic tribes to settle closer to or within Roman-controlled areas, adopting Roman economic practices such as the use of Roman currency and luxury goods. This gradual integration influenced tribal social structures and increased their political complexity.

Everyday life near the limes involved a mix of military and civilian routines, with garrisons, watchtowers, and fortresses serving as foci for both Roman soldiers and local populations. Archaeological evidence points to the coexistence of military installations and agrarian settlements, where locals engaged in farming, livestock raising, and the provisioning of Roman outposts with cereals, cattle, and wood. [46]

Living along the Limes Landscape and settlement in the Lower Rhine Delta during Roman and Early Medieval times

Marieke van Dinter’s dissertation provides a comprehensive analysis of how landscape dynamics and human settlement interacted in the Lower Rhine Delta from the Roman to the Early Medieval periods. The work is notable for its integration of geological, geomorphological, and archaeological data, yielding insights into settlement patterns, military logistics, and landscape use in a challenging deltaic environment. [47]

Van Dinter constructed highly detailed palaeogeographical maps of the Oude Rijn distributary, enabling the study of how landscape changes influenced settlement locations over time.​ These reconstructions clarify the relationship between river dynamics, crevasse splay formation, and human habitation, showing that even short-lived landscape features could provide settlement sites from the Neolithic onward.

Illustration Eighteen: Palaegeographic Map of Netherlands First Century CE

Click for Larger View | Source: Figure 2.2 Palaeogeographic map of the Netherlands in 1st century AD Box indicates the research area, in Marieke van Dinter, Living Along the Limes, PhD Thesis, 23 Aug 1970, Utrecht Studies in Earth Sciences 2017

The dissertation challenges the long-held assumption that Roman military sites along the Rhine were built next to abandoned meanders (oxbow lakes) for safety. Instead, Van Dinter’s research demonstrates that most Roman forts and settlements were established on the raised edges of active meanders, specifically on the undercut banks of the main river channel.

These locations were chosen for their strategic advantages: they allowed for the construction of quays that ensured year-round mooring and access to the river, essential for military logistics and trade.

Contrary to previous beliefs that the Roman army in the Rhine-Meuse delta was supplied mainly from distant sources, Van Dinter’s analysis of archaeological data suggests that local agrarian settlements played a significant role in provisioning the army with wood, cereals, and beef. Modelling of the landscape’s carrying capacity and the army’s supply needs revealed that the local environment could support a much larger population and military presence than previously assumed

The dissertation documents how major environmental changes, such as the avulsion (course change) of the Rhine and increased flooding frequency between 250 and 850 CE, influenced settlement abandonment and relocation, particularly after the withdrawal of the Roman limes.

From the Augustan period, the Lower Rhine–Meuse zone became a permanent military corridor (limes), with chains of forts and roads whose garrisons required large, regular supplies of grain, fodder, timber and other basics.​ Studies of plant remains and rural sites in the Rhine delta show that farms in the immediate limes hinterland intensified cereal production and participated directly in provisioning the army, while more distant loess zones remained closer to mixed subsistence farming. [48]

The Meuse Rhine Watershed Area as a Economic Hub and Networked Periphery to the Roman Empire

The Rhine–Meuse delta functioned as a transport hub linking the North Sea, Britain, and the German Rhineland to the interior via river and canal routes. Reconstructed canals and boat finds show organized inland navigation connecting the limes with the Scheldt and lower Meuse regions.​ Timber, building materials, and other resources were extracted on a large scale for fort construction and road building, while river ports and quays along active Rhine meanders facilitated continuous movement of bulk goods and military supplies. [49]

The demographic character of the limes zone was defined primarily by its role as an area of immigration, commerce and defence, with constant influxes and mixing of populations rather than static, isolated communities. Roman frontier populations underwent constant demographic redistribution and adaptation—migration, workforce management, and regional realignment—not simply the decline or collapse of indigenous groups. [50]

Civilian settlement patterns, household composition, and local labor organization were strongly conditioned by military presence and recruitment cycles, “civil society” demography could not be modelled independently of the army. Recruitment and veteran settlement in the limes zone were structurally redistributive forces. They removed able-bodied men from local communities, re‑concentrated them in military places, and then partially “returned” them as veterans, reshaping the age–sex structure of surrounding populations. [51]

The so-called “Romanization” of local groups was not uniform. Archaeological and anthropological analyses reveal a spectrum of adoption, adaptation, and resistance to Roman material culture, language, burial customs, and settlement organization. In the Batavian and civitas Tungrorum areas, for example, local material culture diversified and hybrid forms emerged, displaying a blend of indigenous traditions and Roman imports. The persistence of native identities and practices is documented well into the Roman period, supporting a nuanced, bottom-up model of culture change. [52]

The Rhine river basin, as a frontier Roman territory, was not economically isolated and not just a part of the periphery of the Roman Empire. Instead it became an engine of regional development and adaptation, grounded in both military investment and local agency. Economic activity along the Rhine was dynamic, with evidence for agricultural intensification, specialization, and commercial production, demonstrating responsiveness to both Roman imperial demands and local environmental constraints. Roman infrastructural and administrative investments in the region catalyzed enduring patterns of connectivity with other areas, which continued to shape economic behavior and regional integration well beyond the end of Roman rule (see illustration nineteen). [53]

Illustration Nineteen: Late Roman Military and Roadway Infrastructure

Click for Larger View |Source: Fig. 1. Late Roman military forces on the frontiers: command centres and fortifications in Raymond Brulet, The Roman army and military defense in Northern Gaul and the Germanic provinces during the Late Empire, Nico Roymans, Stijn Heeren & Wim De Clercq, eds, Social Dynamics in the Northwest Frontiers of the Late Roman Empire, page 36, Amsterdam University Press, 2017 , https://www.researchgate.net/publication/365257551_The_Roman_army_and_military_defence_in_Northern_Gaul_and_the_Germanic_provinces_during_the_Late_Empire

Roman state and military investments led to a dramatic rise in connectivity, integrating coastal areas into a broader imperial context and transforming rural life of indigenous groups. This is aparent in the middle to upper Rhine river areas. The construction of Roman roads, waterways, and quays—most notably the Corbulo Canal and the limes road—greatly increased mobility and integration, particularly from the middle of the second century onward (see illustration nineteen). [54]

The rural communities became increasingly connected to wider networks. Towns like Voorburg were granted municipium status, catalyzing urban development and acting as administrative and economic centers. There was a marked rise in imported goods found at rural sites, indicating intensified regional trade and economic prosperity for parts of the population. Roman-style buildings and material culture proliferated as connectivity increased. Military installations and urban centers played a critical role in facilitating and sustaining connectivity, including interactions with the Roman fleet (Classis Germanica), as evidenced by finds of stamped artifacts and inscriptions (re: illustration nineteen). [55]

These transformations were ultimately subject to broader environmental and political challenges. Many rural settlements were abandoned in the early third century due to rising water tables. By the mid-to-late third century, habitation in both military and urban centers declined sharply, except for continued activity in major Roman cities and maintenance of main roads. The highest degree of connectivity and prosperity was reached in the late second and early third centuries, followed by abrupt decline and depopulation by the late third to early fourth century. While the Roman presence drastically altered local settlement and economic patterns, these changes were not permanent, illustrating the vulnerability of regional prosperity to environmental and imperial shifts. [56]

When reviewing illustration nineteen above, it is apparent that there were fewer roads connecting cities, towns and forts with the rural areas in the lower Rhine / delta area. A study by Philipe Verhagen corroborates this distinction. Verhagen finds that rural settlements in the Dutch Roman border region of the Lower Rhine region exhibited a relatively weak socioeconomic hierarchy compared to urban and military centers. The research highlights limited interaction between towns, forts, and their rural hinterlands (see illustration twenty). Verhagen’s findings, in contrast to the middle and upper portion of the Limes areas in the Lowland, identify a distinctive lack of strong central places in the lower Rhine river area and emphasize minimal economic or social interdependence between towns, forts, and rural communities. [57]

This lack of road network structure in the lower Rhine River area suggests a more fragmented network, shaped more by location of military installations and river logistics than by market-driven or state-imposed centrality. Verhagen’s case shows that while core models of hierarchy and centralization work in some provinces, frontier zones could manifest far weaker forms of settlement network organization. [58]

Illustration Twenty: Map of the Lower Rhine region with major towns and indigenous groups, with the Verhagen study region outlined in red

Click for larger View | Source: Fig 1. in Verhagen, P. Centrality on the periphery: an analysis of rural settlement hierarchy in the Dutch part of the Roman limes. Archaeol Anthropol Sci 15, 45 (2023). https://doi.org/10.1007/s12520-023-01745-0

The Changing Composition of Indigenous Groups

Charles Joseph O’Hara, in his PhD thesis, proposes a multidisciplinary re-evaluation of the concept of an ethnic divide between Celts and Germans during this period. By synthesizing historical, linguistic, and archaeological evidence, O’Hara challenges the traditional boundaries and narratives established by Roman sources. The overall conclusion is that identity and culture in the Rhine and Germania regions during this period were highly fluid, marked by ongoing processes of migration, assimilation, and hybridization rather than fixed ethnic divides. [59]

Greco-Roman writers, starting with Poseidonius and especially Julius Caesar, helped create an artificial divide between “Celts” (Galli) west of the Rhine and “Germans” east of it. Caesar manipulated these definitions for political reasons, solidifying the concept of an ethnic dichotomy that lacked real contemporary tribal self-identification. [60]

The thesis documents how tribal names, personal names, place names, and the archaeological record reveal the hybridization and ethnogenesis of new cultural groups, such as the Chatti, Batavi, Cugerni, Tungri, and Texuandri, reflecting both Celtic and Germanic influences. Material culture in “Germanic” areas is sometimes found to be entirely La Tène (Celtic), demonstrating the fluidity of ethnic and cultural boundaries. Major demographic and cultural shifts—prompted by migration waves from northern and eastern Germania (Teutonic zones) into the Rhine frontier and Central Europe—led to the transformation of the ethnic, linguistic, and material landscape, producing hybrid societies with mixed identities.

Following the settlement of Germanic groups and after Caesar’s Gallic campaigns, there was large-scale movement and expansion of Teutonic tribes from northern and eastern Germania into central Europe. This process led to the fall of traditional Celtic centers (oppida) and the assimilation of Celts into emerging Germanic confederacies, contributing to new “hybrid” ethnic identities such as the Chatti. [61]

By the end of the first century CE, many of the so-called “Germanic” tribes had substantial Celtic roots, having adopted new languages and material cultures due to migrations and political change. The multicultural, multilingual, and hybrid nature of the region’s population is attested both linguistically and archaeologically. Language cannot reliably distinguish Gallic (Celtic) from Germanic identity, nor can material culture alone.

Ethnogenesis in this period was the product of ongoing movement, cultural exchange, and amalgamation of diverse ‘tribal’ societies. O’Hara’s thesis strongly emphasizes integrating evidence from all three disciplines—history, linguistics, and archaeology—to arrive at a reconstruction of early European ethnogenesis, moving beyond the simplified narratives imposed by Roman authors (see table three). [62]

The Roman era catalyzed deep transformations in the composition, distribution, and social structures of both Germanic and Celtic groups, through conquest, migration pressures, and cultural integration that shaped early medieval Europe. [63]

Table Three: Changing Tribal Composition in Roman Era

TribePeriod ActiveApprox. RegionNotes
Germani cisrhenaniLate 2nd–1st c. BCEBetween lower Meuse and RhineBelgic tribes with early Germanic influence
Ubii39 BCE onwardCologne area (West bank Rhine)Voluntarily allied with Rome
Cugerni (Sugambri)8 BCE onwardNear Xanten (Lower Rhine)Resettled east-Rhine tribe
Batavians (Chatti)Late 1st c. BCE–3rd c. CERhine–Meuse deltaNotable Roman auxiliaries
Tencteri & Usipetes1st c. BCEEast of Lower RhineEarly Germanic intruders
Frisians & Chauci1st–3rd c. CERhine–Meuse estuaryNorth Sea Germans expanding south
Franks3rd–5th c. CELower Rhine & ToxandriaDominant post‑Roman group

The Late-Roman Social and Demographic Collapse

The late-Roman social and demographic collapse may have also had an impact on the absence of identified haplogroups of the Griff(is)(es)(ith) paternal line. The social and demographic collapse in the late Roman era occurred between the late fourth and late fifth centuries CE. [64] The collapse spanned roughly a century, with its most acute effects felt between 400 and 476 CE. Internal socio-economic decline was evident as early as the Crisis of the Third Century (c. 235–284 CE), marked by civil wars, epidemics, and economic trouble in the Roman Empire.[65] The demographic and social unraveling accelerated sharply following the crossing of the Rhine by Germanic groups in 406/407, the sack of Rome in 410, and ongoing barbarian invasions and fragmentation.[66]

The process culminated in 476 CE when Romulus Augustulus was deposed, ending central imperial control in the West.​ While some scholars point to continued infrastructure and regional Romanized practices into the early sixth century, the principal period of collapse is broadly agreed to be around 400–476 CE. [67]

The late-Roman social and demographic collapse was due to a complex interplay of factors, including environmental disasters, epidemics, social instability, economic decline, migration pressures, and regional genetic turnover, as revealed by archaeological, genetic, and anthropological research. [68]

Archaeological and climate studies point to climate deterioration, notably the Late Antique Little Ice Age, which led to crop failures and food shortages that triggered famines.​Epidemics, including plagues such as the Antonine Plague and later the Justinian Plague, caused abrupt population declines and eroded social resilience.​ Malnutrition and declining living standards are directly evidenced archaeologically, as average nutrition and health markers decrease in the fourth and fifth centuries. [69]

Economic collapse was reflected in the marked decline of urban centers, reduction in per-capita output, and weakening of imperial taxation and bureaucratic structures, resulting in widespread ruralization and urban abandonment. Social fragmentation intensified as Roman political and social systems failed, leading to the breakdown of order and widened individual hardship and class inequality. [70]

Large-scale migrations occurred as the Roman frontier collapsed under pressure from groups such as Huns, Goths, and other ‘barbarian’ peoples — many of these groups forced to migrate westward by climate changes in Eurasia. [71] Ancient DNA and isotope studies show marked genetic turnover, with new populations (many genetically similar to Slavic-speaking groups in the Balkans or northern Europeans elsewhere) arriving and replacing previous local populations, creating one of Europe’s largest permanent demographic changes. [72] Large population movements within Europe introduced Slavic speakers into areas formerly populated by Germans, Romance speakers into areas formerly populated by Thracians and Dacians, and German speakers in areas formerly populated by Romance speakers. [73]

The major plagues of the Roman era—the Antonine Plague, Plague of Cyprian, and Justinianic Plague—had differing impacts on Belgica, Germania Inferior, and the Rhine–Meuse Delta, with the frontier regions often experiencing less severe or more ambiguous effects than Mediterranean core provinces (see table four).

Table Four: Pandemic Outbreaks and Timing

EventDate RangeImpact/Correlation
Drought, cooling, instability [74] 3rd–4th c.Agricultural crisis, migration
Antonine Plague [75]165–180 CEFirst major pandemic, onset of decline
Plague of Cyprian 250–270 CESecond pandemic, continued losses
“Late Antique Little Ice Age” [76]530s–540s CESevere cooling, crop failure, famine
Justinian Plague [77]541–549 CE & afterMassive mortality, further collapse

The Antonine Plague, thought to be smallpox, was a catastrophic pandemic for the Roman Mediterranean and core provinces. Archaeological and economic survey data from Germania Inferior and neighboring provinces show some demographic decline and contraction in rural settlement after the Antonine Plague but with less dramatic impact recorded than in the urbanized Mediterranean heartland. The effect in regions like Germania Inferior appears in the archaeological record as stagnation or slow decline—rather than mass catastrophe—likely due to lower population density, limited urbanization, and more rural/flexible settlement structures. [79]

The Cyprian Plague’s most visible impact is documented in Roman North Africa, Egypt, and Italy.​ There is little direct archaeological or written evidence that it caused major demographic collapse in the Rhine–Meuse region or Germania Inferior, though indirect effects (military disruption, economic weakening) may have contributed to long-term decline. [80]

The Justinianic Plague devastated the Eastern Roman Empire and parts of the Mediterranean, with catastrophic mortality and economic disruption.​ Recent research questions the extent to which the plague severely affected Frankish Europe and the former western frontier provinces, including Belgica and the Rhine–Meuse delta. Contemporary sources and modern studies suggest the black rat vector (and therefore the disease) was less prevalent in northern and western Europe during the sixth century, so the mortality impact was likely less severe in these areas compared to the cities of the Mediterranean basin.​ Archaeological and genetic evidence from these northern regions indicate some demographic contraction, but not the unprecedented mortality apparent farther south and east; many modern reviews argue that the Justinianic Plague’s reach into the Rhine–Meuse was limited or indirect. [81]

The timing and severity of demographic decline during the late-Roman collapse varied considerably across western provinces, with key regional differences shaped by invasions, environmental change, local resilience, and economic context. In frontier areas such as the Meuse–Rhine watershed area, the collapse was felt chiefly through settlement abandonment, incursions of groups from Germania, population decline, and the end of effective Roman administration, especially after 410 and more drastically after 450–476 CE. [82]

Repeated invasions and migrations by groups from the periphery (often termed “barbarians”) placed immense pressure on Roman borders and internal stability. The breakdown of centralized authority and the fragmentation of political power led to the erosion of administrative and economic systems, weakening the ability to respond to crises.

The late Roman Era Crisis of the third century, the formation of the Gallic Empire, and further settlement and integration of Germanic populations, including forced or negotiated settlement of Frankish groups, are used by authors Roymans, Derks and Heeren as a case study to illustrate the effects of Roman imperial expansion, population movement, and rural reordering during the late Roman era crisis. Their argument is corroborated by archaeological signs of settlement abandonment and later reoccupation. [83]

The historical developments in the fourth and early fifth century in Germania secunda were determined by two key themes. The first is the continuous attempt of the Roman authorities to control the Rhine corridor in order to protect the Gallic hinterland against Germanic raiders and to keep open the strategic routes to Britannia via the Rhine and Meuse.[84]

The second theme . . . , concerns the substantial influx of Germanic groups, or more specifically the groups described as Franks in the written sources, from the period around 400 onwards. Franci is a collective name for a series of smaller tribes in the areas east and north of the Lower Rhine who had long maintained relations with the Roman Empire. Not until the middle of the third century does this name appear in the written accounts. In the third and fourth centuries Franks were generally described as people living outside the Roman Empire, but in the late fourth and fifth centuries they also inhabited land in Germania secunda. Frankish society underwent a major transformation during the late Roman period, which was closely tied to increasing interaction – both friendly and hostile – with the Roman Empire. Viewed from this perspective, the Franks can be regarded as a product of the late Roman frontier. [85]

Ancient DNA studies reveal significant population movements and genetic turnover, especially in frontier regions like the Balkans, where large-scale migrations from Eastern Europe introduced new genetic lineages.​ The collapse of Roman urban centers and the ruralization of populations are reflected in both archaeological and genetic data, with evidence of declining urban populations and increased genetic heterogeneity. [86]

The decline of long-distance trade networks and the collapse of complex economic systems led to a reduction in social stratification and the loss of specialized crafts and professions.​ Archaeological evidence points to a shift from urban to rural living, with the abandonment of cities and a reorganization of settlement patterns.

Anthropological models, such as “barbarigenesis” (see sidebar discussion), suggest that the formation of new societies on the periphery of the Roman Empire was both a cause and consequence of Roman decline, as peripheral groups adapted to and exploited the weaknesses of Roman power.​ The “barbarigenesis” model views the formation of new societies at the Roman Empire’s periphery as a reciprocal process with Roman decline, where peripheral groups developed more power relative to their wealth by adapting to and exploiting Roman weaknesses. [87]

This is seen as a dynamic of social simplification, as the core Roman society declined and peripheral societies adapted, resulting in a decrease in social complexity, especially as wealth and power became more aligned across regions over time. [88]

The Barbarigenesis Model Explaining the collapse of complex societies such as the Roman Empire [89]

The core-periphery dynamic and wealth-power mismatch 

  • Wealth and power imbalance: A key concept in the model is the mismatch between wealth and power. The Roman core focused more on wealth production, while peripheral societies focused more on fighting.
  • Opportunity cost: Peripheral groups could be more successful because the opportunity cost of fighting was lower for them than for the richer, more powerful Roman core.
  • Consequence of the mismatch: This imbalance created a long-lasting decline in social complexity, with the effects spreading from the more developed center to the less developed periphery, as social and political structures simplified over time. 

Social simplification

Another key concept of this model is social simplification.

  • Diminished complexity: The collapse was characterized by social simplification, marked by a reduction in markers of complex society.
  • Decline in infrastructure and literacy: This included a diminished investment in large-scale monumental architecture, such as the grand public buildings, and a decline in the use of literacy across the population.
  • Formation of new social structures: As the Roman Empire fractured, new social structures, like the manorial system and later feudalism, began to develop at the local level to provide protection and manage resources in the absence of a central authority

The late-Roman collapse had tangible impacts on the Meuse–Rhine watershed, transforming its demographic, environmental, and cultural landscape through population decline, settlement abandonment, intensified flooding, and notable genetic shifts. The area is characterized by two major periods of population growth: the middle Roman period (70–270 CE) and the early medieval period C (725–950 CE).​ Between these periods, a significant population decline occurred during the late Roman period (270–450 CE), with population numbers dropping by 78–85 percent. [90]

In a paper “Counting heads: Post-Roman population decline in the Rhine-Meuse delta (the Netherlands) and the need for more evidence-based reconstructions”, authors Rowin J. van Lanen and Bert Groenewoudt present a quantitative method for modeling demographic fluctuations in the Rhine-Meuse delta during the first millennium CE, focusing on the transition from the Roman to the early medieval period. [91]

As identified by van Lanen and Groenewoudt, within the Rhine–Meuse delta, several sub‑regions experienced the late/post‑Roman decline with markedly different timing and severity, especially when comparing the coastal belt, the central river area, and the higher coversand zones inland. Ilustration twenty-one depicts the diverging declines conditioned by local geomorphology, hydrology, and land‑use.

  • Coastal and tidal‑influenced zone (western delta and estuarine margins): Here, the combination of marine influence, peat and tidal landscapes, and high geomorphological instability produced strong settlement contraction and, in some sectors, near‑abandonment in the late Roman and earliest post‑Roman phases. Recovery of rural occupation was relatively late, only becoming substantial again in the later early medieval period.
  • Fluvial levee and central river zone (Rhine and Waal levees and adjacent floodplains): This zone shows the classic “boom–bust–slow recovery” pattern: very high densities in the middle Roman period, a severe drop in the late Roman period (around 80% loss), followed by a prolonged low plateau and then renewed growth from ca. 8th–9th centuries. Within this, some levee segments with more favourable elevation and hydrological conditions retained or regained settlement earlier than others.
  • Inland coversand and higher Pleistocene grounds (eastern and south‑eastern margins of the delta): These higher, relatively flood‑safe areas show less dramatic contraction and, in some models, a smoother, more gradual demographic curve with smaller amplitude of decline. In several of these inland sectors, rural populations remain comparatively stable or recover earlier than in the low‑lying fluvial and coastal tracts.

Illustration Twenty-One: Reconstructed regional population trends in Late Roman Era and Early Medieval Period

Click for Larger View | Source: Bert Groenewoudt and Rowin J van Lanen, Post-Roman population dynamics in the Rhine- Meuse delta (the Netherlands). RURALIA XII Conference, 11th-17th September, Kilkenny (Ireland), https://www.academia.edu/37462618/Diverging_decline_Reconstructing_and_validating_post_Roman_population_trends_AD_0_1000_in_the_Rhine_Meuse_delta_the_Netherlands_?nav_from=15cb6c13-c1d0-4f81-bb5c-0f8de12d669f

The thesis of this study is one of the key studies underpinning the now widely accepted picture of a severe Late Roman–Early Medieval population trough in the Rhine–Meuse delta area, on the order of a 70–80 percent decline from mid‑Roman to late Roman levels in many sectors. The reduced settlement densities and contraction of arable land in the third to fifth centuries led to marked changes in land use intensity, with abandonment of marginal fields and a shift in the balance between cultivated, grazed, and regenerating woodland areas.

A central finding is that post‑Roman woodland regeneration occurred on a large scale in parts of the southern Netherlands and selected sandy landscapes, but was spatially uneven and contingent on earlier land‑use histories. In some eastern and central sandy regions (e.g. Veluwe), palynological records [92] show only modest woodland recovery, whereas in the south extensive reforestation followed the contraction of Roman-period agriculture, implying a mosaic of semi-open to closed woodland rather than a uniformly “re-wilded” landscape. In densely exploited Roman-era core zones, post‑Roman decades saw both demographic contraction and partial ecological “release” (woodland and wetland expansion); whereas other areas show continuity or even renewed intensification in the Early Middle Ages, foreshadowing later medieval reclamation cycles. [93]

Settlements concentrated along high, dry alluvial ridges were abandoned or depopulated; many lower-lying Roman sites became lost due to environmental transformations, as large swathes of land were overtaken by new floodplain processes or buried beneath fluvial sediments.​ A lack of archaeological finds from post-Roman times in key locations implies that occupation patterns shifted, and previously populous riverine and urban centers saw steep reductions in habitation and visibility. [94]

The Genetic Impact of Roman Colonization

Given current data and available research, the safest reading of the genetic impact of Roman colonization is it added a detectable but secondary “cosmopolitan” layer to Y-DNA diversity in the northwestern frontier, superimposed on and later overshadowed by both earlier Bronze Age and later post-Roman demographic events. [95]

The Roman period introduced some new YDNA lineages into the northwestern frontier, but current evidence suggests only modest and uneven impact on the long‑term male-line pool in regions such as Germania Inferior compared with much larger earlier (Bronze Age) and later (Migration/Medieval) shifts. This is inspite of the turbulant and disruptive effects of Roman incursion in the region. Most of the dominant YDNA lineages in the modern Low Countries still appear rooted in pre-Roman (especially Corded Ware / Bell Beaker) and post-Roman (Frankish and later) YDNA lineages rather than in Roman-era soldier or migrant input.

Studies of ancient Y-DNA (aDNA) for the Roman-period Lower Rhine itself is still extremely sparse. This is perhaps due to the impact of Roman incursion and their destructive actions against the indigenous groups, the reordering and mass migration of local groups in the Meuse Rhine waterway region and the lack identification and of testing of human remains in the area.

Most inferences on ancient aDNA distribution rely on pre-Roman aDNA in the Rhine–Meuse region, Roman frontier datasets from other provinces, and later (Merovingian–medieval) and modern Y-chromosome structure in the Low Countries. A 2025 whole‑genome study on the Rhine–Meuse shows early Corded Ware-associated males with R1b-U106 already present by the late third millennium BCE, implying that a key “Germanic” lineage was well established locally long before the Roman conquest. [96]

Roman-era activity on the northwestern frontier appears to have had only a modest, and mostly local, impact on Y‑chromosome lineages: current data point to strong continuity of pre‑Roman “local” haplogroups (especially R1b-U106/R1b-P312 subclades and I1/I2) from the Late Iron Age into the early medieval period, with limited detectable input of distinctly Italian or eastern provincial paternal lines into communities like those of Germani Inferior. The main detectable shifts in YDNA structure in the region seem to cluster around earlier Bronze–Iron Age processes (steppe-derived lineages becoming dominant) and later Migration- and early medieval-period movements, rather than the Roman imperial phase itself. [97]

Holocene aDNA from the Rhine–Meuse region shows that by the later third millennium BCE, male lines had already undergone a major turnover, with Mesolithic I2a/R1b-V88/C1a lineages replaced by steppe-associated Y haplogroups such as R1b-U106 appearing in early Corded Ware contexts. This creates a ‘pre‑Roman baseline’ in which Germanic-speaking Iron Age groups around the Lower Rhine are already dominated by “northern/central European” R1b (U106 and P312-derivatives) and I1/I2 lineages, the same broad clades that continue into historical times. [98]

Genome-wide work on Roman frontiers elsewhere (Danube, Balkans) shows surprisingly little long-term Y‑chromosome impact from Italian colonists and legionaries: typical Italian Iron Age lineages such as R1b-U152 remain rare even in areas with heavy Roman military presence, and local or Balkan lineages (e.g. E-V13, I2, R1b-M269) dominate male burials. This pattern supports a model where recruitment was increasingly local and where foreign soldiers often did not leave enduring male lines in frontier rural communities, a scenario that is consistent with the limited evidence from the northwestern limes. [99]

A study of 348 male individuals from the Netherlands, dated c. 500–1850 CE, found that Y‑chromosome diversity changes across time can be largely explained by drift under a patrilocal regime, with no strong evidence for major paternal discontinuities between early medieval and later populations. This implies that whatever Roman-era demographic perturbations occurred in the wider region, they did not produce a clear, lasting break in Y‑haplogroup composition in the post-Roman Low Countries, which remained dominated by R1b and I lineages throughout. [100]

Synthesizing these lines of evidence, the Roman era in the northwest frontier appears to have:

  • Left the main pre‑existing male lineages (R1b-U106/P312, I1, I2, C1a) largely intact into the post‑Roman period, suggesting continuity of local patrilines across the imperial frontier centuries.
  • Introduced some non-local Y lineages through soldiers, merchants, and slaves (including Mediterranean and possibly African clades), but these are so far detected as rare or idiosyncratic rather than forming new dominant haplogroups.
  • Maintained genetic continuity among local populations, with no large-scale replacement of male lineages, as most settlements absorbed rather than supplanted indigenous groups; despite the influx of new cultural and administrative practices and the exploitative and repressive aspects of Roman rule.
  • Fostered some degree of genetic mixing from Roman military presence and urban development, but the demographic impact was diluted by prevailing local kinship patterns, rural settlement structures, and social boundaries that limited the integration of outside male lineages into the general population.
  • Prepared the social and demographic setting (through urban nodes, mobility, and later military recruitment) for substantial early medieval mixing and migration, whose genetic effects—especially in autosomes and mtDNA—are clearer than any distinct Roman Y‑chromosomal horizon in the Germani Inferior zone.

The net result was a paradoxical pattern: despite the collective social and cultural practices of the Roman Empire, a strong substrate of indigenous male Y-DNA diversity in the Meuse-Rhine watershed was preserved throughout the period. The Lower Rhine Limes, in particular, shows a distinctive genetic landscape with ancient G2a and I2a haplogroups preserved in Roman enclaves and an expansion of R1b-U106 after the Roman collapse.

Recent ancient DNA studies from the medieval Low Countries found that Y-chromosome structure became regionally entrenched in the patrilocal societies of the Middle Ages. Limited male mobility and local authority boundaries reinforced haplogroup differences, helping to preserve and differentiate lineages on a local scale, as seen in the patterns for R1b-U106 (Germanic, north/east of the Rhine) and R1b-P312 (Celtic, south/west), with residual pre-Indo-European (G2a) lineages in Roman and isolated enclaves. [101]

Most of the “absence” of documented haplogroups reflects methodological and contextual limitations of aDNA work on the Roman-period Low Countries rather than a real impossibility of reconstructing phylogenetic trees. Current projects only rarely have the depth, coverage, and sample base needed to build robust, fine-grained Y‑DNA phylogenies specifically for Roman-era lineages in this region.

The Y chromosome is a tiny fraction of total DNA and is especially hard to recover from degraded ancient bone, so shotgun sequencing often yields too few reliable Y‑SNPs to place Roman-period males deeply in a phylogeny. Common capture panels (e.g. 1240k) were designed around modern variation, underrepresent many older or rare lineages, and thus often allow only broad haplogroup calls rather than branch-level trees for local Roman samples. [102]

Ancient Y‑chromosome datasets frequently have low coverage with substantial post‑mortem damage, which provides low statistical confidence ranges for phylogenetic inference. To avoid overinterpreting ‘statistical noise’, many studies restrict themselves to assigning individuals to known haplogroups instead of attempting full lineage trees that would require high-quality sequence across thousands of informative sites. [103]

For the Low Countries, Roman-period skeletons with good DNA preservation are relatively scarce and unevenly distributed compared to early medieval and later burials, so sample sizes by site and horizon are often too small for robust local Y‑phylogenies. Several major Low Countries Y‑chromosome projects explicitly target early medieval to modern periods to study patrilocality and recent substructure, leaving the Roman era poorly represented. [104]

Many Roman sites in the Rhine–Meuse–Scheldt region are military, urban, or cremation-heavy contexts, all of which tend to yield poorer skeletal preservation and fewer suitable male inhumations than later rural cemeteries. Where preservation is good (e.g. some frontier and rural cemeteries), research agendas often focus on overall ancestry and mobility rather than detailed male-line descent, so only coarse Y‑haplogroup assignments are reported. [105]

Genetic drift and population bottlenecks can entirely remove Y‑chromosome lineages over time, and both processes are now seen as major contributors to the loss and reshaping of male lines in Europe, including the Low Countries, especially across the Holocene and into the historical period. In finite populations, random variation in male reproductive success means that some Y lineages increase in frequency while others decline and disappear purely by chance, without any selective advantage.

This effect is amplified in small, fragmented, or patrilocal communities, such as the Lowlands of the Meuse Rhine watershed area, where local male effective population sizes are low and drift can rapidly erase rare or moderately frequent lineages. In highly structured, patrilocal landscapes like much of northwestern Europe after late antiquity, repeated local bottlenecks from war, epidemic, or economic disruption could remove entire village‑level patrilines, cumulatively contributing to the disappearance of many Roman‑period Y lineages even if overall census size recovered later.

Sources:

Feature Banner: The banner at the top of the story features two tribal maps of the Lower Rhine frontier zone in the Caesarian and early Imperial periods. Overlayed on the maps is estimated migratory path of the Griff(is)(es)(ith) paternal YDNA line that is associated with the phylogenetic gap that is the major topic of the multipart story. The maps are from an article by Nico Roymans, Ton Derks, Stijn Heeren, Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative, Britannia, 51, 24 Jun 2020 (1 Nov 2020), 265 – 294, https://research.vu.nl/en/publications/roman-imperialism-and-the-transformation-of-rural-society-in-a-fr/

[1] Mata, Karim, Anthropological Perspectives on Colonialism, Globalisation and Rural Lifeways: Expanding the Limits of Archaeological Interpretation in the Lower Rhineland, Pages: 33–47 in Duggan, M., McIntosh, F. and Rohl, D.J (eds.) (2012) TRAC 2011: Proceedings of the Twenty First Annual Theoretical Roman Archaeology Conference, Newcastle 2011. Oxford: Oxbow Books.

Manuel Fernández-Götz, Dominik Maschek and Nico Roymans, The Dark Side of the Empire: Roman Expansionism between Object Agency and Predatory Regime, https://ora.ox.ac.uk/objects/uuid:772a01a4-e5f4-4094-8cac-563e1e1f7d0f/files/s6t053g09b

Nico Roymans, Ton Derks, Stijn Heeren, Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative, Britannia, 51, 24 Jun 2020 (1 Nov 2020), 265 – 294, https://research.vu.nl/en/publications/roman-imperialism-and-the-transformation-of-rural-society-in-a-fr/

Fernandez-Gotz, M, Maschek, D & Roymans, N 2020, ‘The dark side of the Empire: Roman expansionism between object agency and predatory regime’, Antiquity, vol. 94, no. 378, pp. 1630 – 1639.

https://doi.org/10.15184/aqy.2020.125 , https://www.pure.ed.ac.uk/ws/files/184974216/Fernandez_GotzMA2020TheDarkSide.pdf

Gardner A. Re-balancing the Romans. Antiquity. 2020;94(378):1640-1642. doi: https://10.15184/aqy.2020.170 

Diederick Habermehl, Julie Van Kerckhove, Nico Roymans, Lisette Kootker, Gerard Boreel, Dennis Braekmans, and Stijn Heeren, Investigating migration and mobility in the Early Roman frontier. The case of the Batavi in the Dutch Rhine delta (c. 50/30 BC–AD 40) , Germania 100, 2022, 65-108

[2] See James Griffis, The Griff(is)(es)(ith) Patrilineal Line of Descent: The Shape and Movement of the G Phylogenetic Tree through Time, 23 Mar 2025, Griffis Family Blog, https://griffis.org/the-griffisesith-patrilineal-line-of-descent-the-shape-and-movement-of-the-g-phylogenetic-tree-through-time/

[3] Martin’s research relied on:

  • the interpretation of shifts, breaks, or continuities in material culture and occupation as evidence for changing sociopolitical dynamics, migration, and identity negotiation within and between tribal groups;
  • the typological study of pottery assemblages from numerous excavated sites, enabling chronological phase distinctions and regional cultural characterization;
  • spatial analysis of settlements and artifact distributions to identify recurrent patterns associated with specific cultural groups, using both published and unpublished excavation data as well as literature from regional surveys;
  • the synthesis of archaeological data with ancient written sources (such as Caesar and other Roman authors), using these texts cautiously to cross-check or contextualize archaeological models rather than to define a priori boundaries.

Martin focused on the civitas Tungrorum area, comparing settlement density, site hierarchies (notably oppida, rural settlements, and isolated farms), and the presence of culturally diagnostic artifacts to propose the most likely zones of influence for different tribal entities.

Overall, the mapping derived from a cross-disciplinary framework integrating ceramic typology, settlement archaeology, and judicious historical comparison, with an emphasis on variability and negotiation of tribal identities rather than static boundaries is noteworthy.

See: Martin, F. , Recent research on material culture and territorial dynamics of late Iron Age tribes in the middle Meuse basin, on the eve of the Roman Conquest, Abb. 10 Proposed localization of the main Celtic tribes of the area, 2019, in : Spätlatènezeitliche und frühkaiserzeitliche Archäologie zwischen Maas und Rhein, Bonn, 323-334. https://www.academia.edu/43850047/Martin_F_2019_Recent_research_on_material_culture_and_territorial_dynamics_of_late_Iron_Age_tribes_in_the_middle_Meuse_basin_on_the_eve_of_the_Roman_Conquest_In_Spätlatènezeitliche_und_frühkaiserzeitliche_Archäologie_zwischen_Maas_und_Rhein_Bonn_323_334

[4] Habermehl, D., van Kerckhove, J., Roymans, N., Kootker, L., Boreel, G., Braekmans, D., & Heeren, S. , Investigating migration and mobility in the Early Roman frontier. The case of the Batavi in the Dutch Rhine delta c. 50/30 BC–AD 40). Germania100(1-2), 65-108. 2022, https://research.vu.nl/ws/portalfiles/portal/265709035/Germania_Investigating_migration_and_mobility_in_the_Early_Roman_frontier_The_case_of_the_Batavi_in_the_Dutch_Rhine_delta.pdf

Berber Sanderijn van der Meulen-van der Veen, Cultural Transformations in Germania Secunda A Holistic Approach to ‘Barbarian’ Migrations, Oxfordshire: Araeopress Publishing Ltd, 2025, https://www.archaeopress.com/Archaeopress/DMS/95CA439F52F34DFB96093E2D8E3A8740/9781803279916-sample.pdf

Hofmann D, Burmeister S, Furholt M, Johannsen NN. Ethnicity in European archaeology: revitalizing a concept with the study of migration. Archaeological Dialogues. Published online 2025:1-19. doi:10.1017/S1380203825100081, https://www.cambridge.org/core/journals/archaeological-dialogues/article/ethnicity-in-european-archaeology-revitalizing-a-concept-with-the-study-of-migration/64E6D3CFAE32024C0B343B282C46800C

O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, https://era.ed.ac.uk/handle/1842/29925

[5] Nico Roymans is an Emeritus Professor of West-European Archaeology at the Vrije Universiteit Amsterdam, specializing in the archaeology of the Late Iron Age and the integration of Celtic and Germanic societies into the Roman world. In his published work he emphasizes that processes of tribal ethnogenesis—such as for the Batavians—did not occur in isolation but as responses to the pressures and opportunities of Roman imperial presence. The Romans actively contributed to the shaping of tribal identities at the imperial frontier, sometimes fostering the emergence of more clearly defined and militarized ethnic groups (like the Batavians) that served specific roles, such as supplying auxiliary troops to the Roman army. He highlights how these identities could shift, be re-invented, or gain new significance as local groups negotiated their place within the imperial structure.

Key to Roymans’ perspective is the view that ethnic group unity is ideologically constructed, often involving myth, tradition, and invention, and is open to challenge or redefinition in response to changing political and social circumstances. Collective identities were far from primordial. Instead, the Roman era witnessed dynamic developments where such identities could be reinforced, transformed, or even invented, partly through imperial policies and interactions with the Roman state. Roymans interprets tribal group identity and transformation as fluid, historically contingent processes, with Roman imperialism playing a crucial, sometimes catalytic, role in shaping, promoting, or even creating the ethnic groups recognized in historical and archaeological sources.

Roymans contrasts essentialist and social-constructionist models by critiquing the former as overly rigid and static, while championing the latter as more sensitive to historical dynamics and context. Essentialist models treat ethnic and tribal identities as fixed, primordial, and rooted in inherent biological or cultural traits that persist unchanged over long periods; these perspectives ignore how group identities can change or be reinvented in response to social and political developments.

In contrast, Roymans’s social-constructionist approach emphasizes that tribal and ethnic identities are ideologically constructed, situational, and often involve invented traditions or myths of origin. He stresses that these identities are fluid, contingent on external circumstances such as imperial intervention, and subject to active negotiation, challenge, and even abandonment. Adopting this model, Roymans argues for an understanding of identity as a product of lived experience, power structures, and shifting traditions, rather than as a timeless essence.

See: Nico Roymans, Ethnic Identity and Imperial Power. The Batavians in the Roman Empire. Amsterdam, Archaeological Studies, volume 10. Amsterdam University Press, 2004.

For similar perspectives see also:

Moore, T. , Detribalizing the later prehistoric past: concepts of tribes in Iron Age and Roman studies. Journal of Social Archaeology 11(3): 334-360) , 2011 

Cristina Crizbasan,  Moving Communities and Changing Ceramics: The Impact of Batavian Auxiliaries across the Roman Empire, PhD Thessis, 2025

Habermehl, D., van Kerckhove, J., Roymans, N., Kootker, L., Boreel, G., Braekmans, D., & Heeren, S. , Investigating migration and mobility in the Early Roman frontier. The case of the Batavi in the Dutch Rhine delta c. 50/30 BC–AD 40). Germania100(1-2), 65-108. 2022, https://research.vu.nl/ws/portalfiles/portal/265709035/Germania_Investigating_migration_and_mobility_in_the_Early_Roman_frontier_The_case_of_the_Batavi_in_the_Dutch_Rhine_delta.pdf

O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, https://era.ed.ac.uk/handle/1842/29925

[6] Berber Sanderijn van der Meulen-van der Veen’s “Cultural Transformations in Germania Secunda” delivers a nuanced, data-driven reassessment of the Lower German frontier’s in the Late Roman era transformations. The research reinterprets frontier change in general by looking at local socio-economic processes and the continuity of critical material culture.

Van der Meulen-van der Veen argues that population change was gradual, shaped by long-term depopulation and resettlement fueled by regional interactions, not abrupt ethnic replacement. Van der Meulen-van der Veen stresses that construction techniques and much of the material culture can reflect adaptation and local living patterns. Van der Meulen-van der Veen uses settlement pattern archaeology, metallurgical studies, and landscape evidence to reconstruct multi-phase occupation and long-term transformation.

See:

Berber Sanderijn van der Meulen-van der Veen, Cultural Transformations in Germania SecundaA Holistic Approach to ‘Barbarian’ Migrations, Oxfordshire: Araeopress Publishing Ltd, 2025, https://www.archaeopress.com/Archaeopress/DMS/95CA439F52F34DFB96093E2D8E3A8740/9781803279916-sample.pdf

In general, see:

Hofmann D, Burmeister S, Furholt M, Johannsen NN. Ethnicity in European archaeology: revitalizing a concept with the study of migration. Archaeological Dialogues. Published online 2025:1-19. doi:10.1017/S1380203825100081, https://www.cambridge.org/core/journals/archaeological-dialogues/article/ethnicity-in-european-archaeology-revitalizing-a-concept-with-the-study-of-migration/64E6D3CFAE32024C0B343B282C46800C

[7] Hofmann D, Burmeister S, Furholt M, Johannsen NN. Ethnicity in European archaeology: revitalizing a concept with the study of migration. Archaeological Dialogues. Published online 2025:1-19. doi:10.1017/S1380203825100081, https://www.cambridge.org/core/journals/archaeological-dialogues/article/ethnicity-in-european-archaeology-revitalizing-a-concept-with-the-study-of-migration/64E6D3CFAE32024C0B343B282C46800C

Berber Sanderijn van der Meulen-van der Veen, Cultural Transformations in Germania SecundaA Holistic Approach to ‘Barbarian’ Migrations, Oxfordshire: Araeopress Publishing Ltd, 2025, https://www.archaeopress.com/Archaeopress/DMS/95CA439F52F34DFB96093E2D8E3A8740/9781803279916-sample.pdf

[8] Hofmann D, Burmeister S, Furholt M, Johannsen NN. Ethnicity in European archaeology: revitalizing a concept with the study of migration. Archaeological Dialogues. Published online 2025:1-19. doi:10.1017/S1380203825100081, https://www.cambridge.org/core/journals/archaeological-dialogues/article/ethnicity-in-european-archaeology-revitalizing-a-concept-with-the-study-of-migration/64E6D3CFAE32024C0B343B282C46800C

[9] Hofmann D, Burmeister S, Furholt M, Johannsen NN. Ethnicity in European archaeology: revitalizing a concept with the study of migration. Archaeological Dialogues. Published online 2025:1-19. doi:10.1017/S1380203825100081, https://www.cambridge.org/core/journals/archaeological-dialogues/article/ethnicity-in-european-archaeology-revitalizing-a-concept-with-the-study-of-migration/64E6D3CFAE32024C0B343B282C46800C

[10] Habermehl, D., van Kerckhove, J., Roymans, N., Kootker, L., Boreel, G., Braekmans, D., & Heeren, S. , Investigating migration and mobility in the Early Roman frontier. The case of the Batavi in the Dutch Rhine delta c. 50/30 BC–AD 40). Germania100(1-2), 65-108, 2022, https://www.academia.edu/106603054/Investigating_migration_and_mobility_in_the_Early_Roman_frontier_The_case_of_the_Batavi_in_the_Dutch_Rhine_delta_c_50_30_BC_AD_40_

Hofmann D, Burmeister S, Furholt M, Johannsen NN. Ethnicity in European archaeology: revitalizing a concept with the study of migration. Archaeological Dialogues. Published online 2025:1-19. doi:10.1017/S1380203825100081, https://www.cambridge.org/core/journals/archaeological-dialogues/article/ethnicity-in-european-archaeology-revitalizing-a-concept-with-the-study-of-migration/64E6D3CFAE32024C0B343B282C46800C

[11] O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, https://era.ed.ac.uk/handle/1842/29925

Batavi (Germanic tribe), Wikipedia, This page was last edited on 27 October 2025, https://en.wikipedia.org/wiki/Batavi_(Germanic_tribe)

[12] O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, https://era.ed.ac.uk/handle/1842/29925

Roymans, Nico, Ethnic Identity and Imperial Power : The Batavians in the Early Roman Empire, Amsterdam: Amsterdam University Press, 2004, https://www.academia.edu/56940817/Ethnic_Identity_and_Imperial_Power_The_Batavians_in_the_Early_Roman_Empire

Wetzels, Eric , Terra Mosana Storyline Language changes in the Euregio Meuse Rhine between 100 BC and 1000 AD DEF, Workpackage 2 – Deliverable No. DT 2.2.1 (A) Language,

[13] See, for example:

Sneath, David. (2016) 2023. “Tribe”. In The Open Encyclopedia of Anthropology, edited by Felix Stein. Facsimile of the first edition in The Cambridge Encyclopedia of Anthropology. Online: http://doi.org/10.29164/16tribe

Moore, T. (2011). Detribalizing the later prehistoric past: concepts of tribes in Iron Age and Roman studies. Journal of Social Archaeology, 11(3), 334-360. https://doi.org/10.1177/1469605311403861; https://durham-repository.worktribe.com/output/1534900/detribalizing-the-later-prehistoric-past-concepts-of-tribes-in-iron-age-and-roman-studies

Fried, Morton H., The Notion of Tribe, Menlo Park: Morton Fried’s 1975 book, 1975, https://www.scribd.com/doc/51384839/Morton-Fried-The-Notion-of-Tribe

Derks, A. M. J. (2009). Ethnic identity in the Roman frontier. The epigraphy of Batavi and other Lower Rhine tribes. In A. M. J. Derks, & N. G. A. M. Roymans (Eds.), Ethnic constructs in antiquity. The role of power andtradition (pp. 239-276). (Amsterdam Archaeological Studies; No. 13). Amsterdam University Press. https://research.vu.nl/ws/portalfiles/portal/2406626/DerksEthnicity.pdf

[14] Sneath, David. (2016) 2023. “Tribe”. In The Open Encyclopedia of Anthropology, edited by Felix Stein. Facsimile of the first edition in The Cambridge Encyclopedia of Anthropology. Online: http://doi.org/10.29164/16tribe

O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, https://era.ed.ac.uk/handle/1842/29925

Fried, Morton H., The Notion of Tribe, Menlo Park: Morton Fried’s 1975 book, 1975, https://www.scribd.com/doc/51384839/Morton-Fried-The-Notion-of-Tribe

[15] Nienhuis, Piet H., Environmental History of the Rhine–Meuse Delta, Springer Science + Business Media B.V., 2008, Page 17, https://www.researchgate.net/profile/Piet-Nienhuis/publication/242024465_Environmental_History_of_the_Rhine-Meuse_Delta_An_Ecological_Story_on_Evolving_Human-Environmental_Relations_Coping_with_Climate_Change_and_Sea-level_Rise/links/53e7d1090cf2fb74872386e6/Environmental-History-of-the-Rhine-Meuse-Delta-An-Ecological-Story-on-Evolving-Human-Environmental-Relations-Coping-with-Climate-Change-and-Sea-level-Rise.pdf

[16] Sneath, David. (2016) 2023. “Tribe”. In The Open Encyclopedia of Anthropology, edited by Felix Stein. Facsimile of the first edition in The Cambridge Encyclopedia of Anthropology. Online: http://doi.org/10.29164/16tribe

O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, https://era.ed.ac.uk/handle/1842/29925

Fried, Morton H., The Notion of Tribe, Menlo Park: Morton Fried’s 1975 book, 1975, https://www.scribd.com/doc/51384839/Morton-Fried-The-Notion-of-Tribe

[17] Commentarii de Bello Gallico, Wikipedia, This page was last edited on 30 October 2025, https://en.wikipedia.org/wiki/Commentarii_de_Bello_Gallico

Caesar, Julius, Commentarii de Bello Gallico, with notes by George Long,  London, Whittaker, 1871, https://archive.org/details/commentariid00caes/page/n3/mode/2up

Germania (book), Wikipedia, This page was last edited on 21 October 2025, https://en.wikipedia.org/wiki/Germania_(book)

Sheposh, Richard, Germanic Peoples, 2024, EBSCO Knowledge Advantage, https://www.ebsco.com/research-starters/military-history-and-science/germanic-peoples

Rhineland, Wikipedia, This page was last edited on 5 November 2025,  https://en.wikipedia.org/wiki/Rhineland

List of ancient Celtic peoples and tribes, Wikipedia, This page was last edited on 20 October 2025, https://en.wikipedia.org/wiki/List_of_ancient_Celtic_peoples_and_tribes

List of early Germanic peoples, Wikipedia, This page was last edited on 1 November 2025, https://en.wikipedia.org/wiki/List_of_early_Germanic_peoples

[18] Hofmann D, Burmeister S, Furholt M, Johannsen NN. Ethnicity in European archaeology: revitalizing a concept with the study of migration. Archaeological Dialogues. Published online 2025:1-19. doi:10.1017/S1380203825100081

[19] Sneath, David. (2016) 2023. “Tribe”. In The Open Encyclopedia of Anthropology, edited by Felix Stein. Facsimile of the first edition in The Cambridge Encyclopedia of Anthropology. Online: http://doi.org/10.29164/16tribe

[20] See the following sources:

Eburones, Wikipedia, This page was last edited on 14 October 2025, https://en.wikipedia.org/wiki/Eburones

Atuatuci, Wikipedia, This page was last edited on 20 April 2025, https://en.wikipedia.org/wiki/Atuatuci

Nervii, Wikipedia, This page was last edited on 30 October 2025, https://en.wikipedia.org/wiki/Nervii

Segri, Wikipedia, This page was last edited on 15 April 2024, https://en.wikipedia.org/wiki/Segni_(tribe)

Treveri, Wikipedia, This page was last edited on 1 August 2025, https://en.wikipedia.org/wiki/Treveri

Condrusi, Wikipedia, This page was last edited on 23 January 2025, https://en.wikipedia.org/wiki/Condrusi

Caerosi, Wikipedia, Wikipedia, This page was last edited on 23 August 2025, https://en.wikipedia.org/wiki/Caerosi

Caerosi, Jatland, This page was last edited on 3 October 2022, https://www.jatland.com/home/Caerosi

Menapii, Wikipedia, This page was last edited on 10 September 2025, https://en.wikipedia.org/wiki/Menapii

[21] Eburones, Jatland, This page was last edited on 12 October 2022, https://www.jatland.com/home/Eburones

[22] Lendering, Jona, Eburones, Last Modified on 5 August 2020, Livius, https://www.livius.org/articles/people/eburones/

See also Toorians, Laura, “Aduatuca, ‘place of the prophet’. The names of the Eburones as representatives of a Celtic language, with an excursus on Tungri”, in: Creemers, G. (ed.), Archaeological contributions to materials and immateriality, Atvatvca 4 (2013) 108-121, https://www.academia.edu/33460316/Toorians_Aduatuca_pdf

[23] Eburones, Wikipedia, This page was last edited on 14 October 2025, https://en.wikipedia.org/wiki/Eburones

[24] Eburones, Jatland, This page was last edited on 12 October 2022, https://www.jatland.com/home/Eburones

[25] Potter, David, ‘Rebellion and Reconstruction’, Master of Rome: A Life of Julius Caesar (New York , 2025; online edn, Oxford Academic, 24 July 2025), 170-194 https://doi.org/10.1093/oso/9780190867188.003.0013

Roymans, Nico & Fernandez-Gotz, Manuel, Caesar in Gaul: New perspectives on the archaeology of mass violence in T. Brindle, M. Allen, E. Durham & A . Smith (eds) TRAC 2014: Proceedings of the Twenty-Four Theoretical Roman Archaelogy Conference, Oxbow Books, 2015,pp 70-80, www.ooxbowbooks.com/oxbow/trac-2014.html , https://www.pure.ed.ac.uk/ws/portalfiles/portal/22191489/2015_Caesar_in_Gaul._New_perspectives_on.pdf

Eburones, Wikipedia, This page was last edited on 14 October 2025, https://en.wikipedia.org/wiki/Eburones

[26] Potter, David, ‘Rebellion and Reconstruction’, Master of Rome: A Life of Julius Caesar (New York , 2025; online edn, Oxford Academic, 24 July 2025), 170-194 https://doi.org/10.1093/oso/9780190867188.003.0013

Roymans, Nico & Fernandez-Gotz, Manuel, Caesar in Gaul: New perspectives on the archaeology of mass violence in T. Brindle, M. Allen, E. Durham & A . Smith (eds) TRAC 2014: Proceedings of the Twenty-Four Theoretical Roman Archaelogy Conference, Oxbow Books, 2015,pp 70-80, www.ooxbowbooks.com/oxbow/trac-2014.html , https://www.pure.ed.ac.uk/ws/portalfiles/portal/22191489/2015_Caesar_in_Gaul._New_perspectives_on.pdf

Eburones, Wikipedia, This page was last edited on 14 October 2025, https://en.wikipedia.org/wiki/Eburones

[27] Eburones, Wikipedia, This page was last edited on 14 October 2025, https://en.wikipedia.org/wiki/Eburones

Gallic Wars, Wikipedia, This page was last edited on 2 November 2025, https://en.wikipedia.org/wiki/Gallic_Wars

[28] Lendering, Jona, Eburones, Last Modified on 5 August 2020, Livius, https://www.livius.org/articles/people/eburones/

Eburones, Wikipedia, This page was last edited on 14 October 2025, https://en.wikipedia.org/wiki/Eburones

Roymans, Nico & Fernandez-Gotz, Manuel, Caesar in Gaul: New perspectives on the archaeology of mass violence in T. Brindle, M. Allen, E. Durham & A . Smith (eds) TRAC 2014: Proceedings of the Twenty-Four Theoretical Roman Archaelogy Conference, Oxbow Books, 2015,pp 70-80, www.ooxbowbooks.com/oxbow/trac-2014.html , https://www.pure.ed.ac.uk/ws/portalfiles/portal/22191489/2015_Caesar_in_Gaul._New_perspectives_on.pdf

Tungri, Wikipedia, This page was last edited on 7 August 2025, https://en.wikipedia.org/wiki/Tungri

[29] See for example:

Manuel Fernández-Götz, Dominik Maschek and Nico Roymans, The Dark Side of the Empire: Roman Expansionism between Object Agency and Predatory Regime, https://ora.ox.ac.uk/objects/uuid:772a01a4-e5f4-4094-8cac-563e1e1f7d0f/files/s6t053g09b

Urbanus, Jason, Caesar’s Diplomatic Breakdown, Mar/Arp 2016, Archaeology Magazine, https://archaeology.org/issues/march-april-2016/digs-discoveries/trenches-netherlands-roman-battle/

Roymans, Nico & Fernandez-Gotz, Manuel, Caesar in Gaul: New perspective on the archaeology of mass violence in T Brindle, M. Allen, E. Durham & A. Smith  (eds) TRAC 2014: Proceedings of the Twenty-Fourth Theoretical Roman Conference, Oxbrow Books, Oxford, 2015, 70-80, https://www.pure.ed.ac.uk/ws/portalfiles/portal/22191489/2015_Caesar_in_Gaul._New_perspectives_on.pdf

Taylor, Michael, Book Review of Julius Caesar’s Battle for Gaul: New Archaeological Perspectives, Jan 2021 (125.1), American Journal of Archaeology, https://ajaonline.org/book-review/4225/

Schadee, H. 2008: ‘Caesar’s construction of northern Europe: inquiry, contact and corruption in De Bello Gallico’, The Classical Quarterly 58, 158–80, https://scholar.google.com/scholar_lookup?title=Caesar%27s+construction+of+northern+Europe%3A+inquiry%2C+contact+and+corruption+in+De+Bello+Gallico&author=Schadee+H.&publication+year=2008&journal=The+Classical+Quarterly&volume=58&doi=10.1017%2FS0009838808000128

Slofstra, J. 2002: ‘Batavians and Romans on the Lower Rhine: the romanisation of a frontier area’, Archaeological Dialogues 9, 16–3, https://scholar.google.com/scholar_lookup?title=Batavians+and+Romans+on+the+Lower+Rhine%3A+the+romanisation+of+a+frontier+area&author=Slofstra+J.&publication+year=2002&journal=Archaeological+Dialogues&volume=9&doi=10.1017%2FS1380203800002014&pages=16-38

[30] Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294. doi:10.1017/S0068113X20000148, https://www.cambridge.org/core/journals/britannia/article/roman-imperialism-and-the-transformation-of-rural-society-in-a-frontier-province-diversifying-the-narrative/BF7FA75997E14A1B7BA4241E5DC981C3

For related articles, see:

Roymans, N. and Fernández-Götz, M., (2015): Fire and Sword. The archaeology of Caesar’s Gallic War. Military History Monthly 56: 52-56. https://www.academia.edu/12866878/Fire_and_Sword_The_archaeology_of_Caesar_s_Gallic_War

Fernandez-Gotz, Manuel, Identity and Power: The transformation of Iron Age societies in northeast Gaul. Amsterdam Archaeological Studies 21. Amsterdam University Press, Amsterdam, 2014

Roymans, Nico, Conquest, mass violence and ethnic stereotyping: investigating Caesar’s actions in the Germanic frontier zone, Journal of Roman Archaeology, Vol 32, 2019, Page 443, https://doi.org/10.1017/S1047759419000229, https://www.academia.edu/43467135/2019_Conquest_mass_violence_and_ethnic_stereotyping_investigating_Caesar_s_actions_in_the_Germanic_frontier_zone

Riggsby, Andrew, Caesar in Gaul and Rome War in Words, Astin, University of Texaas Press, 2006

[31] Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294. doi:10.1017/S0068113X20000148, https://www.cambridge.org/core/journals/britannia/article/roman-imperialism-and-the-transformation-of-rural-society-in-a-frontier-province-diversifying-the-narrative/BF7FA75997E14A1B7BA4241E5DC981C3

Roymans, Nico, Conquest, mass violence and ethnic stereotyping: investigating Caesar’s actions in the Germanic frontier zone, Journal of Roman Archaeology, Vol 32, 2019, Page 443, https://doi.org/10.1017/S1047759419000229, https://www.academia.edu/43467135/2019_Conquest_mass_violence_and_ethnic_stereotyping_investigating_Caesar_s_actions_in_the_Germanic_frontier_zone

[32] Quote from Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294.

See also:

Roymans, N. 2019: ‘Conquest, mass violence and ethnic stereotyping: investigating Caesar’s actions in the Germanic frontier zone’, Journal of Roman Archaeology 32, 443

Roymans, N., Heeren, S., and De Clercq, W. (eds) 2017: Social Dynamics in the Northwest Frontiers of the Late Roman Empire: Beyond Decline or Transformation, Amsterdam Archaeological Studies 26, Amsterdam, https://scholar.google.com/scholar_lookup?title=Social+Dynamics+in+the+Northwest+Frontiers+of+the+Late+Roman+Empire%3A+Beyond+Decline+or+Transformation&author=Roymans+N.&author=Heeren+S.&author=De+Clercq+W.&publication+year=2017

[33] Quote from Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294.

[34] Civitas, Wikipedia, This page was last edited on 4 November 2025, https://en.wikipedia.org/wiki/Civitas

de Bruin , Jasper, Border communities at the Edge of the Roman Empire. Processes of Change in the Civitas Cananefatium, Amsterdam Archaeological Studies 28. Amsterdam: Amsterdam University Press. 2019, https://www.academia.edu/42688371/Border_communities_at_the_Edge_of_the_Roman_Empire_Processes_of_Change_in_the_Civitas_Cananefatium

[35] Quote from Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294

[36] Quote from Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294

[37] Quote from Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294

[38] Roymans, Nico and Stijn Heeren, Introduction. New perspectives on the Late Roman Northwest, page 5, in Nico Roymans, Stijn Heeren & Wim De Clercq, eds, Social Dynamics in the Northwest Frontiers of the Late Roman Empire, page 36, Amsterdam University Press, 2017

[39] How Rome Conquered the Ancient Celts, The Roma Empire, https://roman-empire.net/army/how-rome-conquered-the-ancient-celts

Lee, Shawn, Celtic Romanization: Cultural Assimilation or Celtic Romanization: Cultural Assimilation or Cultural Exchange?, 28 Apr 2015, Young Historians Conference, 6, Portland State University, https://pdxscholar.library.pdx.edu/cgi/viewcontent.cgi?article=1071&context=younghistorians

Ancient Celtic Warfare, Wikipedia, This page was last edited on 20 October 2025, https://en.wikipedia.org/wiki/Ancient_Celtic_warfare

Celts, Wikipedia, This page was last edited on 28 September 2025, https://en.wikipedia.org/wiki/Celts

[40] Kooistra, Laura I., The Provisioning of the Roman Army in the Rhine Delta between C. AD 40 and 140, International Congress of Roman Frontier Studies (20th : 2006 : León, Spain). Limes XX Vol. III. Madrid :Ediciones Polifemo : Consejo Superior de Investigaciones Científicas, Instituto Histórico Hoffmeyer, Instituto de Arqueología de Merida, 2009. https://www.ancientportsantiques.com/wp-content/uploads/Documents/PLACES/UK-EUNorth/NLlimes-Kooistra2009.pdf

Laura I. Kooistra; Marieke van Dinter; Monica K. Dütting; Pauline van Rijn; Chiara Cavallo: Could the local population of the Lower Rhine delta supply the Roman army? Part 1: The archaeological and historical framework, Journal of Archaeology in the Low Countries 4-2 (April 2013), https://jalc.nl/cgi/t/text/get-pdf6920.pdf?idno=m0402a01;c=jalc

Gerlach R, Meurers-Balke J, and Kalis AJ. The Lower Rhine (Germany) in Late Antiquity: a time of dissolving structures. Netherlands Journal of Geosciences, Volume 101, e14. https://doi.org/10.1017/njg.2022.11

van Lanen RJ, de Kleijn MTM, Gouw-Bouman MTIJ, Pierik HJ. Exploring Roman and early-medieval habitation of the Rhine–Meuse delta: modelling large-scale demographic changes and corresponding land-use impact. Netherlands Journal of Geosciences. 2018;97(1-2):45-68. doi:10.1017/njg.2018.3

Cavallo, Chiara and , Laura I. Kooistra and Monica K. Düttingin, Food supply to the Roman army in the Rhine delta in the First Century A.D., in Sue Stallibrass and Richard Thomas, eds, Feeding the Roman Army the Archaeology of Production and Supply in NW Europe, Oxford: Oxbow Books, 2008, 69-82, https://www.academia.edu/6359653/Cavallo_et_al_2008_Food_supply_to_the_Roman_army_in_the_Rhine_delta_in_the_first_century_A_D

Thomas, Richard and Sue Stallibrass, For starters: producing and supplying food to the army in the Roman north-west provinces, in Sue Stallibrass and Richard Thomas, eds, Feeding the Roman Army the Archaeology of Production and Supply in NW Europe, Oxford: Oxbow Books, 2008, 1 – 18, https://www.academia.edu/6359653/Cavallo_et_al_2008_Food_supply_to_the_Roman_army_in_the_Rhine_delta_in_the_first_century_A_D

[41] van Lanen RJ, de Kleijn MTM, Gouw-Bouman MTIJ, Pierik HJ. Exploring Roman and early-medieval habitation of the Rhine–Meuse delta: modelling large-scale demographic changes and corresponding land-use impact. Netherlands Journal of Geosciences. 2018;97(1-2):45-68. doi:10.1017/njg.2018.3 , https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/exploring-roman-and-earlymedieval-habitation-of-the-rhinemeuse-delta-modelling-largescale-demographic-changes-and-corresponding-landuse-impact/40F68343AEEC8FF41124C5F098069863

de Soto, P., Pažout, A., Brughmans, T. et al. Itiner-e: A high-resolution dataset of roads of the Roman Empire. Sci Data 12, 1731 (2025). https://doi.org/10.1038/s41597-025-06140-z

Derks, Ton, Town-country dynamics in Roman Gual. The epigraphy of the ruling elite, in Nico Roymans& Ton Derks, Villa Landscapes in the Roman North, Amsterdam, Amsterdam University Press, 2011

Roymans, Ethnic recruitment, returning veterans and the diffusion of Roman culture among rural populations in the Rhineland frontier zone, in in Nico Roymans& Ton Derks, Villa Landscapes in the Roman North, Amsterdam, Amsterdam University Press, 2011

Nico Roymans and Marianne Zandstra, Indications for rural slavery in the northern provinces, in in Nico Roymans & Ton Derks, Villa Landscapes in the Roman North, Amsterdam, Amsterdam University Press, 2011

Dinter, Marieke, The Roman Limes in the Netherlands: How a delta landscape determined the location of the military structures, 1 Apr 2013, Geologie en Mijnbouw/Netherlands Journal of Geosciences, 92, 11-32, DO – 10.1017/S0016774600000251, https://www.researchgate.net/publication/287527529_The_Roman_Limes_in_the_Netherlands_How_a_delta_landscape_determined_the_location_of_the_military_structures/citation/download

Franconi, T.V. (2021). The Environmental Imperialism of the Roman Empire in Northwestern Europe. In: Erdkamp, P., Manning, J.G., Verboven, K. (eds) Climate Change and Ancient Societies in Europe and the Near East. Palgrave Studies in Ancient Economies. Palgrave Macmillan, Cham. https://doi.org/10.1007/978-3-030-81103-7_11

David J. BreezeAndreas ThielSarah RothThomas Becker, Frontiers of the Roman Empire: The Upper Germanic Limes, Oxford: Archaepress Publishing, 2022 , https://www.archaeopress.com/Archaeopress/download/9781803271743

Krogstad , Peter, Limes Germanicus – Barrier or Highway? The Transfer of Technology between the Roman World and Northwestern Europe during the Iron Age – A Maritime Perspective, , https://www.academia.edu/2515458/Limes_Germanicus_Barrier_or_Highway_The_Transfer_of_Technology_between_the_Roman_World_and_Northwestern_Europe_during_the_Iron_Age_A_Maritime_Perspective

Berber van der Meulen-van der Veen, The Late Roman limes revisited. The changing function of the Roman army in the Dutch river/coastal area (AD 260-406/7), Masters Thesis, 18 Jun 2017, https://www.academia.edu/35391375/The_Late_Roman_limes_revisited_The_changing_function_of_the_Roman_army_in_the_Dutch_river_coastal_area_AD_260_406_7_?nav_from=9a71d3d7-3906-4b8e-a8a2-f7b086734a27

[42] Roymans, N. 2019: ‘Conquest, mass violence and ethnic stereotyping: investigating Caesar’s actions in the Germanic frontier zone’, Journal of Roman Archaeology 32, 443

Battle of the Teutoburg Forest, Wkipedia, This page was last edited on 25 October 2025, https://en.wikipedia.org/wiki/Battle_of_the_Teutoburg_Forest

[43] Huntink, Emmelie, Roman Limes in the Rhine Delta: A historical perspective on the early developments of the north-western frontier of the Roman Empire, MA Thesis Ancient History, Leiden: Universiteit Leiden, 2022, https://studenttheses.universiteitleiden.nl/access/item%3A3446973/view

Frontiers of the Roman Empire – The Lower German Limes, UNESCO, https://whc.unesco.org/en/list/1631/

Renger de Bruin, Astrid Hertog and Roeland Paardekooper, The Roman Limes on the Lower Rhine: A European Border’s Visibility in Landscape and Museums, Museum international, 2, 2018, 116-125 , https://www.icom-italia.org/wp-content/uploads/2018/02/ICOMItalia.MuseumInternational.Articolo.RengerdeBruin.AstridHertog.RoelandPaardekooper.pdf

Dinter, Marieke, The Roman Limes in the Netherlands: How a delta landscape determined the location of the military structures, 1 Apr 2013, Geologie en Mijnbouw/Netherlands Journal of Geosciences, 92, 11-32, DO – 10.1017/S0016774600000251, https://www.researchgate.net/publication/287527529_The_Roman_Limes_in_the_Netherlands_How_a_delta_landscape_determined_the_location_of_the_military_structures/citation/download

Franconi, T.V. (2021). The Environmental Imperialism of the Roman Empire in Northwestern Europe. In: Erdkamp, P., Manning, J.G., Verboven, K. (eds) Climate Change and Ancient Societies in Europe and the Near East. Palgrave Studies in Ancient Economies. Palgrave Macmillan, Cham. https://doi.org/10.1007/978-3-030-81103-7_11

David J. BreezeAndreas ThielSarah RothThomas Becker, Frontiers of the Roman Empire: The Upper Germanic Limes, Oxford: Archaepress Publishing, 2022 , https://www.archaeopress.com/Archaeopress/download/9781803271743

Krogstad , Peter, Limes Germanicus – Barrier or Highway? The Transfer of Technology between the Roman World and Northwestern Europe during the Iron Age – A Maritime Perspective, , https://www.academia.edu/2515458/Limes_Germanicus_Barrier_or_Highway_The_Transfer_of_Technology_between_the_Roman_World_and_Northwestern_Europe_during_the_Iron_Age_A_Maritime_Perspective

Berber van der Meulen-van der Veen, The Late Roman limes revisited. The changing function of the Roman army in the Dutch river/coastal area (AD 260-406/7), Masters Thesis, 18 Jun 2017, https://www.academia.edu/35391375/The_Late_Roman_limes_revisited_The_changing_function_of_the_Roman_army_in_the_Dutch_river_coastal_area_AD_260_406_7_?nav_from=9a71d3d7-3906-4b8e-a8a2-f7b086734a27

[44] “Packwerk” structures at Xanten are ancient Roman bank protection constructions, identified through archaeological excavation at the site of the former Roman city, 
Colonia Ulpia Traiana. These structures were built to stabilize the riverbank, and a well-preserved example was found with square timber posts and piles made of soil. They provide evidence of sophisticated engineering used by the Romans to manage the river’s course. 

Gerlach R, Meurers-Balke J, and Kalis AJ. The Lower Rhine (Germany) in Late Antiquity: a time of dissolving structures. Netherlands Journal of Geosciences, Volume 101, e14. https://doi.org/10.1017/njg.2022.11

See related work on the LImes:

Lines (Roman Empire), Wikipedia, This page was last edited on 20 November 2025, https://en.wikipedia.org/wiki/Limes_(Roman_Empire)

Frontiers of the Roman Empire, UNESCO, not dated, https://whc.unesco.org/en/list/430/

Lendering, Jona, Limes, last modified on 10 October 2020, Livius.org, https://www.livius.org/articles/concept/limes/

Krogstad , Peter, Limes Germanicus – Barrier or Highway? The Transfer of Technology between the Roman World and Northwestern Europe during the Iron Age – A Maritime Perspective, , https://www.academia.edu/2515458/Limes_Germanicus_Barrier_or_Highway_The_Transfer_of_Technology_between_the_Roman_World_and_Northwestern_Europe_during_the_Iron_Age_A_Maritime_Perspective

[45] See for example:

Marieke van Dinter, Living Along the Limes, PhD Thesis, 23 Aug 1970, Utrecht Studies in Earth Sciences 2017, https://www.academia.edu/35123998/Living_along_the_Limes_Landscape_and_settlement_in_the_Lower_Rhine_Delta_during_Roman_and_Early_Medieval_times

Brunner, Katrin, The Emperor’s new frontier, modified 10 Jul 2025,Blog Schweizerisches Naticnal Museum, https://blog.nationalmuseum.ch/en/2021/04/roman-frontier/

Harry van Enckevort, Mark Driessen, Erik Graafstal, Tom Hazenberg, Tatiana Ivleva and Carol van Driel-Murray, eds, Living and Dying on the Roman Frontier and Beyond (Limes XXV Volume 3), Leiden: Sidestone Press, 2024 , https://www.sidestone.com/books/living-and-dying-on-the-roman-frontier-and-beyond-limes-xxv-volume-3

Collins, Rob, and Frances McIntosh, editors. Life in the Limes: Studies of the People and Objects of the Roman Frontiers. Oxbow Books, 2014. JSTOR, https://doi.org/10.2307/j.ctvh1dk1v

Dinter, Marieke, The Roman Limes in the Netherlands: How a delta landscape determined the location of the military structures, 1 Apr 2013, Geologie en Mijnbouw/Netherlands Journal of Geosciences, 92, 11-32, DO – 10.1017/S0016774600000251, https://www.researchgate.net/publication/287527529_The_Roman_Limes_in_the_Netherlands_How_a_delta_landscape_determined_the_location_of_the_military_structures/citation/download

[46] Tomas, Angieszka, Living with the Army I. Civil Settlements near Roman Legionary Fortresses in Lower Moesia, Warsaw: University of Warsaw, 2017

Tomas. Angieszka, Life on the Frontier: Roman Military Families in Lower Moesia,  Studia Europaea Gnesnensia, 16 2016, 225-257, https://www.academia.edu/35662809/LIFE_ON_THE_FRONTIER_ROMAN_MILITARY_FAMILIES_IN_LOWER_MOESIA?nav_from=0aa553b7-73ac-4941-b487-cb5c9eb620ee

Lendering, Jona, Germania Inferior (7), last modified on 10 October 2020, Livius.org, https://www.livius.org/articles/place/germania-inferior/germania-inferior-7/

Marieke van Dinter, Living Along the Limes, PhD Thesis, 23 Aug 1970, Utrecht Studies in Earth Sciences 2017, https://www.academia.edu/35123998/Living_along_the_Limes_Landscape_and_settlement_in_the_Lower_Rhine_Delta_during_Roman_and_Early_Medieval_times

[47] Marieke van Dinter, Living Along the Limes, PhD Thesis, 23 Aug 1970, Utrecht Studies in Earth Sciences 2017, https://www.academia.edu/35123998/Living_along_the_Limes_Landscape_and_settlement_in_the_Lower_Rhine_Delta_during_Roman_and_Early_Medieval_time

[48] Ployer, René and Marinus Polak & Ricarda Schmidt, The Frontiers of the Roman Empire, Munich 2017, https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/exploring-roman-and-earlymedieval-habitation-of-the-rhinemeuse-delta-modelling-largescale-demographic-changes-and-corresponding-landuse-impact/40F68343AEEC8FF41124C5F098069863

Marieke van Dinter, Living Along the Limes, PhD Thesis, 23 Aug 1970, Utrecht Studies in Earth Sciences 2017, https://www.academia.edu/35123998/Living_along_the_Limes_Landscape_and_settlement_in_the_Lower_Rhine_Delta_during_Roman_and_Early_Medieval_times

[49] Gerlach R, Meurers-Balke J, and Kalis AJ. The Lower Rhine (Germany) in Late Antiquity: a time of dissolving structures. Netherlands Journal of Geosciences, Volume 101, e14. https://doi.org/10.1017/njg.2022.11

[50] See for example: Isabelle Séguy, Current Trends in Roman Demography and Empirical Approaches to the Dynamics of the Limes Populations, Séguy, Isabelle. (2019). Current Trends in Roman Demography and Empirical Approaches to the Dynamics of the Limes Populations, in: Verhagen, P., Joyce, J., Groenhuijzen, M.R. (eds) Finding the Limits of the Limes. Computational Social Sciences, Springer, Cham. https://doi.org/10.1007/978-3-030-04576-0_2 also https://www.academia.edu/85930389/Current_Trends_in_Roman_Demography_and_Empirical_Approaches_to_the_Dynamics_of_the_Limes_Populations?nav_from=1e1f77cc-1ab0-44ee-8298-379e3f5b9e7c

[51] Séguy, Isabelle, Current Trends in Roman Demography and Empirical Approaches to the Dynamics of the Limes Populations, Séguy, Isabelle. (2019). Current Trends in Roman Demography and Empirical Approaches to the Dynamics of the Limes Populations

Verhagen, Philip and Jamie Joyce, Mark Groenhuijzen, Modelling the Dynamics of Demography in the Dutch Roman Limes Zone, LAC2014 Proceedings, Vrije Universiteit Amsterdam, 2016, https://doi.org/10.5463/LAC.2014.62 , https://osf.io/e5frp/files/a8vhc

[52] See for example:

Heeren Stijn, The Material Culture of Small Rural Settlements in the Batavian Area: A Case Study on Discrepant Experience, Creolisation, Romanisation or Globalisation? Pages: 159–173 in Platts, H., Pearce, J., Barron, C., Lundock, J., and Yoo, J. (eds) 2014. TRAC 2013: Proceedings of the Twenty-Third Annual Theoretical Roman Archaeology Conference, King’s College, London. Oxford: Oxbow, 2013

[53] Franconi, Tyler V., Economic Development of the Rhine River Basin in the Roman Period 30 BC – AD 406, PhD Thesis, Oxford University,, Trinity Term 2014, https://www.academia.edu/7670831/_2014_The_Economic_Development_of_the_Rhine_River_Basin_in_the_Roman_Period_30_BC_AD_406_?nav_from=51d94d35-b05a-4ef9-bcd7-1853da5eddb3

Flückiger, Matthias and Hornung, Erik and Larch, Mario and Ludwig, Markus and Mees, Allard, Roman Transport Network Connectivity and Economic Integration (2019). CESifo Working Paper No. 7740, Available at SSRN: https://ssrn.com/abstract=3424723 or http://dx.doi.org/10.2139/ssrn.3424723

Flückiger, Matthias and Erik Hornung, Mario Larch, Markus Ludwig, and Allard Mees, Roman Transport Network Connectivity and Economic Integration, January 2021, https://cmr.uni-koeln.de/sites/cmr/pdf/Hornung_Erik/FHLLM_January2021.pdf

Huntink, Emmelie, Roman Limes in the Rhine Delta: A historical perspective on the early developments of the north-western frontier of the Roman Empire, 2022, Masters Thesis, Universiteit Leiden, https://studenttheses.universiteitleiden.nl/access/item:3446973/view

de Soto, P., Pažout, A., Brughmans, T. et al. Itiner-e: A high-resolution dataset of roads of the Roman Empire. Sci Data 12, 1731 (2025). https://doi.org/10.1038/s41597-025-06140-z

[54] See for example:

de Bruin, Jasper, Connectivity in the south-western part of the Netherlands during the Roman period (AD 0-350), in Corrie Bakels ad Hans Kammermans, eds, Analecta Praehistorica Leidensia 43/44 : 145-157, 2012, https://www.academia.edu/30199226/Connectivity_in_the_south_western_part_of_the_Netherlands_during_the_Roman_period_AD_0_350_

de Bruin , Jasper, Border Communities at the Edge of the Roman Empire. Processes of Change in the Civitas Cananefatium, Amsterdam Archaeological Studies 28. Amsterdam: Amsterdam University Press. 2019, https://www.academia.edu/42688371/Border_communities_at_the_Edge_of_the_Roman_Empire_Processes_of_Change_in_the_Civitas_Cananefatium

[55] de Bruin, Jasper, Connectivity in the south-western part of the Netherlands during the Roman period (AD 0-350), in Corrie Bakels ad Hans Kammermans, eds, Analecta Praehistorica Leidensia 43/44 : 145-157, 2012, https://www.academia.edu/30199226/Connectivity_in_the_south_western_part_of_the_Netherlands_during_the_Roman_period_AD_0_350_

de Bruin , Jasper, Border Communities at the Edge of the Roman Empire. Processes of Change in the Civitas Cananefatium, Amsterdam Archaeological Studies 28. Amsterdam: Amsterdam University Press. 2019

[56] de Bruin, Jasper, Connectivity in the south-western part of the Netherlands during the Roman period (AD 0-350), in Corrie Bakels ad Hans Kammermans, eds, Analecta Praehistorica Leidensia 43/44 : 145-157, 2012

de Bruin , Jasper, Border Communities at the Edge of the Roman Empire. Processes of Change in the Civitas Cananefatium, Amsterdam Archaeological Studies 28. Amsterdam: Amsterdam University Press. 2019

[57] Verhagen, P. Centrality on the periphery: an analysis of rural settlement hierarchy in the Dutch part of the Roman limes. Archaeol Anthropol Sci 15, 45 (2023). https://doi.org/10.1007/s12520-023-01745-0

[58] Verhagen, P. Centrality on the periphery: an analysis of rural settlement hierarchy in the Dutch part of the Roman limes. Archaeol Anthropol Sci 15, 45 (2023). https://doi.org/10.1007/s12520-023-01745-0

[59] O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, https://era.ed.ac.uk/handle/1842/29925

[60] O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, Pages 5, 6, 72, 74, 75  https://era.ed.ac.uk/handle/1842/29925

[61] O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, Pages 5, 6, 72, 74, 75  https://era.ed.ac.uk/handle/1842/29925

[62] O’Hara, Charles Joseph, Celts and Germans of the First Century BC – Second century AD: An Old Question, A modern Synthesis, PhD Thesis, University of Edinburgh. 2005, Pages 5, 6, 72, 74, 75  https://era.ed.ac.uk/handle/1842/29925

[63] See for example

How Rome Conquered the Ancient Celts, The Roma Empire, https://roman-empire.net/army/how-rome-conquered-the-ancient-celts

Lee, Shawn, Celtic Romanization: Cultural Assimilation or Celtic Romanization: Cultural Assimilation or Cultural Exchange?, 28 Apr 2015, Young Historians Conference, 6, Portland State University, https://pdxscholar.library.pdx.edu/cgi/viewcontent.cgi?article=1071&context=younghistorians

Ancient Celtic Warfare, Wikipedia, This page was last edited on 20 October 2025, https://en.wikipedia.org/wiki/Ancient_Celtic_warfare

Celts, Wikipedia, This page was last edited on 28 September 2025, https://en.wikipedia.org/wiki/Celts

[64] Most scholars marking the watershed period from the death of Theodosius I in 395 to the deposition of the last Western Emperor Romulus Augustulus in 476 CE.

See:

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025, https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

Anastasi, Luciano, Timeline of the Fall of Rome: Western Roman Empire (235–476 AD), 22 Dec 2022, Medieval History, https://historymedieval.com/timeline-fall-of-rome/

Harper, Kyle. The fate of Rome. Climate, disease, and the end of an empire. ISBN 978-0-691-19206-2. Princeton University Press 2017

[65]The Crisis of the Third Century, also known as the Military Anarchy or the Imperial Crisis, was a period in Roman history during which the Roman Empire nearly collapsed under the combined pressure of repeated foreign invasions, civil wars and economic disintegration.

Crisis of the Third Century, Wikipedia, This page was last edited on 15 November 2025, https://en.wikipedia.org/wiki/Crisis_of_the_Third_Century

[66] Graetzer, Daniel G., Fall of Rome, 2022, EBSCO Knowledge Advantage, https://www.ebsco.com/research-starters/military-history-and-science/fall-rome

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025, https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

[67] Harper, Kyle. The fate of Rome. Climate, disease, and the end of an empire. ISBN 978-0-691-19206-2. Princeton University Press 2017

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025, https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

[68] White, Robyn, Ramifications of Roman Empire’s Fall Explained by Geneticists, 7 Dec 2013, Newsweek, https://www.newsweek.com/geneticists-new-effects-fall-roman-empire-1850412

Sánchez, Jorge Pisa, Is this the real reason the Roman Empire collapsed?, 8 Aug, 2025, National Geographic, https://www.nationalgeographic.com/history/article/fall-of-ancient-roman-empire?loggedin=true&rnd=1763782034011

Antonio ML, Gao Z, Moots HM, Lucci M, Candilio F, Sawyer S, Oberreiter V, Calderon D, Devitofranceschi K, Aikens RC, Aneli S, Bartoli F, Bedini A, Cheronet O, Cotter DJ, Fernandes DM, Gasperetti G, Grifoni R, Guidi A, La Pastina F, Loreti E, Manacorda D, Matullo G, Morretta S, Nava A, Fiocchi Nicolai V, Nomi F, Pavolini C, Pentiricci M, Pergola P, Piranomonte M, Schmidt R, Spinola G, Sperduti A, Rubini M, Bondioli L, Coppa A, Pinhasi R, Pritchard JK. Ancient Rome: A genetic crossroads of Europe and the Mediterranean. Science. 2019 Nov 8;366(6466):708-714. doi: 10.1126/science.aay6826. PMID: 31699931; PMCID: PMC7093155., (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC7093155/

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025, https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

[69] See for example:

Bowes, Kim. “When Kuznets Went to Rome: Roman Economic Well-Being and the Reframing of Roman History.” Capitalism: A Journal of History and Economics, vol. 2 no. 1, 2021, p. 7-40. Project MUSEhttps://dx.doi.org/10.1353/cap.2021.0000

Sánchez, Jorge Pisa, Is this the real reason the Roman Empire collapsed? , 8 Aug 2025, National Geographic, https://www.nationalgeographic.com/history/article/fall-of-ancient-roman-empire

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025, https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

Little Ice age, Wikipedia, This page was last edited on 24 November 2025, https://en.wikipedia.org/wiki/Little_Ice_Age

Heinz Wanner, Christian Pfister, Raphael Neukom, The variable European Little Ice Age, Quaternary Science Reviews, Volume 287, 2022, 107531, ISSN 0277-3791,
https://doi.org/10.1016/j.quascirev.2022.107531 .
(https://www.sciencedirect.com/science/article/pii/S0277379122001627 )

Antonine Plague, Wikipedia, This page was last edited on 22 November 2025, https://en.wikipedia.org/wiki/Antonine_Plague

Bruun, Christer, “The Antonine Plague and the ‘Third-Century Crisis'”, in Olivier Hekster, Gerda de Kleijn, Danielle Slootjes (ed.), Crises and the Roman Empire: Proceedings of the Seventh Workshop of the International Network Impact of Empire, Nijmegen, June 20–24, 2006. Leiden/Boston: Brill, 2007 (Impact of Empire, 7), 201–218.

Plague of Justinian, Wikipedia, This page was last edited on 23 November 2025, https://en.wikipedia.org/wiki/Plague_of_Justinian

J. Preiser-Kapeller, W. A. McGrath, R. Pfister, Sh. Gong, M. Keller, T. P. Newfield, U. Schamiloglu, U. Büntgen, M. A. Spyrou, B. Averbuch, F. Chen, N. Schindel, H. Xie, E. Xoplaki, The Circulation of Yersinia pestis in Central Eurasia before and during the First Plague Pandemic (Second to Eighth Century CE): Palaeogenetic and Historical Evidence and Sociopolitical, Ecological, and Climatic Factors. Human Ecology 2025
DOI: 10.1007/s10745-025-00617-6

[70] Sánchez, Jorge Pisa, Is this the real reason the Roman Empire collapsed? , 8 Aug 2025, National Geographic, https://www.nationalgeographic.com/history/article/fall-of-ancient-roman-empire

Britanica Editors, Height and decline of imperial Rome, 12 Nov 2025, Britannica, https://www.britannica.com/place/Roman-Empire/Height-and-decline-of-imperial-Rome

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025,  https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

Harper, Kyle. The fate of Rome. Climate, disease, and the end of an empire. ISBN 978-0-691-19206-2. Princeton University Press 2017

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025, https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

Bowes, Kim. “When Kuznets Went to Rome: Roman Economic Well-Being and the Reframing of Roman History.” Capitalism: A Journal of History and Economics, vol. 2 no. 1, 2021, p. 7-40. Project MUSEhttps://dx.doi.org/10.1353/cap.2021.0000

Sánchez, Jorge Pisa, Is this the real reason the Roman Empire collapsed? , 8 Aug 2025, National Geographic, https://www.nationalgeographic.com/history/article/fall-of-ancient-roman-empire

[71] Iñigo Olalde, Pablo Carrión, Ilija Mikić, Nadin Rohland, Swapan Mallick, Iosif Lazaridis, Matthew Mah, Miomir Korać, Snežana Golubović, Sofija Petković, Nataša Miladinović-Radmilović, Dragana Vulović, Timka Alihodžić, Abigail Ash, Miriam Baeta, Juraj Bartík, Željka Bedić, Maja Bilić, Clive Bonsall, Maja Bunčić, Domagoj Bužanić, Mario Carić, Lea Čataj, Mirna Cvetko, Ivan Drnić, Anita Dugonjić, Ana Đukić, Ksenija Đukić, Zdeněk Farkaš, Pavol Jelínek, Marija Jovanovic, Iva Kaić, Hrvoje Kalafatić, Marijana Krmpotić, Siniša Krznar, Tino Leleković, Marian M. de Pancorbo, Vinka Matijević, Branka Milošević Zakić, Anna J. Osterholtz, Julianne M. Paige, Dinko Tresić Pavičić, Zrinka Premužić, Petra Rajić Šikanjić, Anita Rapan Papeša, Lujana Paraman, Mirjana Sanader, Ivana Radovanović, Mirjana Roksandic, Alena Šefčáková, Sofia Stefanović, Maria Teschler-Nicola, Domagoj Tončinić, Brina Zagorc, Kim Callan, Francesca Candilio, Olivia Cheronet, Daniel Fernandes, Aisling Kearns, Ann Marie Lawson, Kirsten Mandl, Anna Wagner, Fatma Zalzala, Anna Zettl, Željko Tomanović, Dušan Keckarević, Mario Novak, Kyle Harper, Michael McCormick, Ron Pinhasi, Miodrag Grbić, Carles Lalueza-Fox, David Reich, A genetic history of the Balkans from Roman frontier to Slavic migrations, Cell, Volume 186, Issue 25, 2023, Pages 5472-5485.e9, ISSN 0092-8674, https://doi.org/10.1016/j.cell.2023.10.018 .
https://www.sciencedirect.com/science/article/pii/S0092867423011352

[72] White, Robyne, Ramifications of Roman Empire’s Fall Explained by Geneticists, 7 Dec 2023, Newsweek, https://www.newsweek.com/geneticists-new-effects-fall-roman-empire-1850412

Antonio ML, Gao Z, Moots HM, Lucci M, Candilio F, Sawyer S, Oberreiter V, Calderon D, Devitofranceschi K, Aikens RC, Aneli S, Bartoli F, Bedini A, Cheronet O, Cotter DJ, Fernandes DM, Gasperetti G, Grifoni R, Guidi A, La Pastina F, Loreti E, Manacorda D, Matullo G, Morretta S, Nava A, Fiocchi Nicolai V, Nomi F, Pavolini C, Pentiricci M, Pergola P, Piranomonte M, Schmidt R, Spinola G, Sperduti A, Rubini M, Bondioli L, Coppa A, Pinhasi R, Pritchard JK. Ancient Rome: A genetic crossroads of Europe and the Mediterranean. Science. 2019 Nov 8;366(6466):708-714. doi: 10.1126/science.aay6826. PMID: 31699931; PMCID: PMC7093155. (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC7093155/

Iñigo Olalde, et al, A genetic history of the Balkans from Roman frontier to Slavic migrations, Cell, Volume 186, Issue 25, 2023, Pages 5472-5485.e9, ISSN 0092-8674,
https://doi.org/10.1016/j.cell.2023.10.018

[73] Drake, B. Changes in North Atlantic Oscillation drove Population Migrations and the Collapse of the Western Roman Empire. Sci Rep 7, 1227 (2017). https://doi.org/10.1038/s41598-017-01289-z

Iñigo Olalde, et al, A genetic history of the Balkans from Roman frontier to Slavic migrations, Cell, Volume 186, Issue 25, 2023, Pages 5472-5485.e9, ISSN 0092-8674,
https://doi.org/10.1016/j.cell.2023.10.018

[74] Drake, B. Changes in North Atlantic Oscillation drove Population Migrations and the Collapse of the Western Roman Empire. Sci Rep 7, 1227 (2017). https://doi.org/10.1038/s41598-017-01289-z

[75] Antonine Plague, Wikipedia, This page was last edited on 29 November 2025,  https://en.wikipedia.org/wiki/Antonine_Plague

[76] Harper, Kyle, How Climate Change and Plague Helped Bring Down the Roman Empire, 19 Dec 2017, Smithsonian Magazine, https://www.smithsonianmag.com/science-nature/how-climate-change-and-disease-helped-fall-rome-180967591/

Metcalfe, Tom, The Roman Empire’s Worst Plagues Were Linked to Climate Change, 26 Jan 2024, Scientific American, https://www.scientificamerican.com/article/the-roman-empires-worst-plagues-were-linked-to-climate-change/

[77] Mordechai, L and M. Eisenberg, T.P. Newfield, A. Izdebski, J.E. Kay, H. Poinar,  The Justinianic Plague: An inconsequential pandemic?, Proc. Natl. Acad. Sci. U.S.A. 116 (51) 25546-25554, https://doi.org/10.1073/pnas.1903797116 (2019).

Harper, Kyle, How Climate Change and Plague Helped Bring Down the Roman Empire, 19 Dec 2017, Smithsonian Magazine, https://www.smithsonianmag.com/science-nature/how-climate-change-and-disease-helped-fall-rome-180967591/

[79] Jongman, W. M., Jacobs, J. P. A. M., & Klein Goldewijk, G. M. (2019). Health and wealth in the Roman Empire. Economics & Human Biology, 34 , 138-150. https://doi.org/10.1016/j.ehb.2019.01.005

[80] Plague of Cyprian, Wikipedia, This page was last edited on 17 November 2025, https://en.wikipedia.org/wiki/Plague_of_Cyprian

[81] Faure, Eric, Did the Justinianic Plague Truly Reach Frankish Europe around 543 AD?,  Vox Patrum 78 (2021) 427-466 DOI: 10.31743/vp.12278, https://10.31743/vp.12278

Latham, Andrew, Justinian’s Plague and the Birth of the Medieval World, Medievalist.net, https://www.medievalists.net/2020/11/justinian-plague-medieval-world/

[82] Demography of the Roman Empire. Wikipedia, This page was last edited on 27 October 2025, https://en.wikipedia.org/wiki/Demography_of_the_Roman_Empire

Baker, David C. , Demographic-Structural Theory and the Roman Dominate, in Leonid E. Gri- nin and Andrey V. Korotayev, eds, History & Mathematics: Trends and Cycles, ‛Uchitel’ Publishing House, 2014, 159 – 189, https://www.academia.edu/24888636/Trends_and_Cycles , also https://www.sociostudies.org/almanac/articles/files/ham/hm_4/pdf/159-189.pdf

Drake, B. Changes in North Atlantic Oscillation drove Population Migrations and the Collapse of the Western Roman Empire. Sci Rep 7, 1227 (2017). https://doi.org/10.1038/s41598-017-01289-z

Lanen, Rowin J.  and Bert Groenewoudt, Counting heads: Post-Roman population decline in the Rhine-Meuse delta (the Netherlands) and the need for more evidence-based reconstructions, in Niall Brady & Claudia Theune, eds, Settlement Change Across Medieval Europe Old Paradigms and New Vistas, 2019, Ruralia XII, 113- 134, https://www.academia.edu/40523376/Counting_heads_Post_Roman_population_decline_in_the_Rhine_Meuse_delta_the_Netherlands_and_the_need_for_more_evidence_based_reconstructions

[83] Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294

See also:

Nico Roymans and Stijn Heeren, Romano-Frankish interaction in the Lower Rhine frontier zone from the late 3rd to the 5th century – Some key archaeological trends explored, Germania99, 2021, 133-156

[84] Quote from Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294

[85] Quote from Roymans N, Derks T, Heeren S. Roman Imperialism and the Transformation of Rural Society in a Frontier Province: Diversifying the Narrative. Britannia. 2020;51:265-294

Romanization (cultural), Wikipedia, This page was last edited on 27 October 2025,  https://en.wikipedia.org/wiki/Romanization_(cultural)

[86] White, Robyn, Ramifications of Roman Empire’s Fall Explained by Geneticists, 7 Dec 2023, Newsweek, https://www.newsweek.com/geneticists-new-effects-fall-roman-empire-1850412

Antonio ML, Gao Z, Moots HM, Lucci M, Candilio F, Sawyer S, Oberreiter V, Calderon D, Devitofranceschi K, Aikens RC, Aneli S, Bartoli F, Bedini A, Cheronet O, Cotter DJ, Fernandes DM, Gasperetti G, Grifoni R, Guidi A, La Pastina F, Loreti E, Manacorda D, Matullo G, Morretta S, Nava A, Fiocchi Nicolai V, Nomi F, Pavolini C, Pentiricci M, Pergola P, Piranomonte M, Schmidt R, Spinola G, Sperduti A, Rubini M, Bondioli L, Coppa A, Pinhasi R, Pritchard JK. Ancient Rome: A genetic crossroads of Europe and the Mediterranean. Science. 2019 Nov 8;366(6466):708-714. doi: 10.1126/science.aay6826. PMID: 31699931; PMCID: PMC7093155. (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC7093155/

Iñigo Olalde, Pablo Carrión, Ilija Mikić, Nadin Rohland, Swapan Mallick, Iosif Lazaridis, Matthew Mah, Miomir Korać, Snežana Golubović, Sofija Petković, Nataša Miladinović-Radmilović, Dragana Vulović, Timka Alihodžić, Abigail Ash, Miriam Baeta, Juraj Bartík, Željka Bedić, Maja Bilić, Clive Bonsall, Maja Bunčić, Domagoj Bužanić, Mario Carić, Lea Čataj, Mirna Cvetko, Ivan Drnić, Anita Dugonjić, Ana Đukić, Ksenija Đukić, Zdeněk Farkaš, Pavol Jelínek, Marija Jovanovic, Iva Kaić, Hrvoje Kalafatić, Marijana Krmpotić, Siniša Krznar, Tino Leleković, Marian M. de Pancorbo, Vinka Matijević, Branka Milošević Zakić, Anna J. Osterholtz, Julianne M. Paige, Dinko Tresić Pavičić, Zrinka Premužić, Petra Rajić Šikanjić, Anita Rapan Papeša, Lujana Paraman, Mirjana Sanader, Ivana Radovanović, Mirjana Roksandic, Alena Šefčáková, Sofia Stefanović, Maria Teschler-Nicola, Domagoj Tončinić, Brina Zagorc, Kim Callan, Francesca Candilio, Olivia Cheronet, Daniel Fernandes, Aisling Kearns, Ann Marie Lawson, Kirsten Mandl, Anna Wagner, Fatma Zalzala, Anna Zettl, Željko Tomanović, Dušan Keckarević, Mario Novak, Kyle Harper, Michael McCormick, Ron Pinhasi, Miodrag Grbić, Carles Lalueza-Fox, David Reich, A genetic history of the Balkans from Roman frontier to Slavic migrations, Cell, Volume 186, Issue 25, 2023, Pages 5472-5485.e9, IISSN 0092-8674, https://doi.org/10.1016/j.cell.2023.10.018 .
(https://www.sciencedirect.com/science/article/pii/S0092867423011352 )

Fall of the Western Roman Empire, Wikipedia, This page was last edited on 9 November 2025, https://en.wikipedia.org/wiki/Fall_of_the_Western_Roman_Empire

[87] The clearest explicit use of a “process in barbarigenesis” model for the Roman-era collapse is the 2021 PLOS One article “Barbarigenesis and the collapse of complex societies: Rome and after” by Eric C. Jones and colleagues, which formulates and tests a quantitative model of barbarian-group formation around the Roman Empire from ca. 1–1200 CE. [a]

While not always using the term “barbarigenesis,” a number of ethnogenesis-oriented scholars analyze the fall of the West as the outcome of long-term processes of group formation at the frontier and within the imperial system:

Reinhard Wenskus, Stammesbildung und Verfassung. Das Werden der frühmittelalterlichen gentes (1961), was foundational for the notion that early medieval “tribes” (gentes) were historically formed political-ethnic groups rather than primordial peoples. This framework is repeatedly cited in discussions of late Roman frontier groups and the emergence of successor kingdoms.[b]

Peter Heather, especially The Fall of the Roman Empire and Invasion of the Barbarians, emphasizes how long-term military and political interaction with Rome fostered increasingly powerful, cohesive barbarian polities whose cumulative pressure contributed decisively to western collapse, explicitly engaging with and partially revising Wenskus-style ethnogenesis. [c]

Early-medieval identity studies influenced by the “ethnogenesis school” (e.g. work discussed in overviews of barbarian identity and ethnogenesis) often treat the post‑Roman kingdoms as the product of ‘negotiated identity formation’ between Roman provincial populations and incoming or militarized “barbarian” elites, aligning conceptually with the barbarigenesis model even where the term is not used.

[87a] Jones, Doug, Barbarigenesis and the collapse of complex societies: Rome and after, 16 Sep 2021, e0254240, PLOS ONE, 16, DO – 10.1371/journal.pone.0254240,
https://www.researchgate.net/publication/354651939_Barbarigenesis_and_the_collapse_of_complex_societies_Rome_and_after

[87b] At the time of writing this story, there is no full English translation of Wenskus’ Stammesbildung und Verfassung. Das Werden der frühmittelalterlichen gentes (1961) in print. The monograph has remained available only in the original German. Its influence in Anglophone scholarship has come via summaries and adaptations, especially in Herwig Wolfram’s works and in secondary literature on ethnogenesis.

For an accessible route into Wenskus’ model (Traditionskern, gentes-formation, etc.), Anglophone scholars typically recommend reading Wolfram’s translated works (e.g. History of the GothsThe Roman Empire and Its Germanic Peoples) and later overviews and critiques that summarize Wenskus’ arguments. There are also English-language articles and book chapters that reconstruct his key theses and situate them within later debates on ethnogenesis and barbarian identity, but these are secondary expositions rather than translations.

Drawing upon studies on modern tribes, Wenskus posited that the Germanic tribes of antiquity did not constitute distinct ethnicities, but were rather diverse alliances led by a dominant elite continuing “core-traditions” (Traditionskerne). Wenskus argued that members of the Germanic tribes were not necessarily related to each other by kin, but rather believed themselves to be.

Vienna School of History, Wikipedia, This page was last edited on 27 May 2025, https://en.wikipedia.org/wiki/Vienna_School_of_History

[87c] Heather traces the Fall of the Western Roman Empire to external migration triggered by the Huns in the late 4th century.

Heather, Peter (2018). “Race, Migration And National Origins”History, Memory and Public Life. Routledge. pp. 80–100.

Bowers, Hannah, “It was Barbarians!” – Peter Heather and Rome’s Decline, 18 Nov 2012, coffeeshopthinking, https://coffeeshopthinking.wordpress.com/2012/11/18/it-was-the-barbarians-the-fall-of-rome-according-to-peter-heather/

[88] Jones, Doug, Barbarigenesis and the collapse of complex societies: Rome and after, 16 Sep 2021, e0254240, PLOS ONE, 16, DO – 10.1371/journal.pone.0254240,
https://www.researchgate.net/publication/354651939_Barbarigenesis_and_the_collapse_of_complex_societies_Rome_and_after

Tainter, Joeseph, The Collapse of Complex Societies, Cambridge, Cambrige University Press, 1988, https://sackett.net/The-Collapse-of-Complex-Societies.pdf

[89] Jones, Doug, Barbarigenesis and the collapse of complex societies: Rome and after, 16 Sep 2021, e0254240, PLOS ONE, 16, DO – 10.1371/journal.pone.0254240,
https://www.researchgate.net/publication/354651939_Barbarigenesis_and_the_collapse_of_complex_societies_Rome_and_after

[90] Rowin J van Lanen and Bert Groenewoudt, Counting heads: Post-Roman population decline in the Rhine-Meuse delta (the Netherlands) and the need for more evidence-based reconstructions, in Niall Brady & Claudia Theune, eds, Settlement Change Across Medieval Europe: Old paradigms and New Vistas, , 2019, Ruralia XII, Leiden; Sidestone, 2019, https://www.academia.edu/40523376/Counting_heads_Post_Roman_population_decline_in_the_Rhine_Meuse_delta_the_Netherlands_and_the_need_for_more_evidence_based_reconstructions

[91] Rowin J van Lanen and Bert Groenewoudt, Counting heads: Post-Roman population decline in the Rhine-Meuse delta (the Netherlands) and the need for more evidence-based reconstructions, in Niall Brady & Claudia Theune, eds, Settlement Change Across Medieval Europe: Old paradigms and New Vistas, , 2019, Ruralia XII, Leiden; Sidestone, 2019, https://www.academia.edu/40523376/Counting_heads_Post_Roman_population_decline_in_the_Rhine_Meuse_delta_the_Netherlands_and_the_need_for_more_evidence_based_reconstructions

[92] Palynological records are archives of past life, made of microscopic fossils like pollen, spores, and dinocysts, that provide clues about ancient environments and climate. By studying these microscopic “palynomorphs” in sediment layers, scientists can reconstruct past vegetation, track climate changes over long periods, determine the age of rocks, and even understand the impact of human activity on ecosystems. 

See:

Palynology, Wikipedia, This page was last edited on 4 October 2025, https://en.wikipedia.org/wiki/Palynology

[93] See for example:

Groenewoudt, B., & Spek, T. , Woodland Dynamics as a Result of Settlement Relocation on Pleistocene Sandy Soils in the Netherlands (200 BC–AD 1400). Rural Landscapes, 3 (1), 2015, 1-17. https://doi.org/10.16993/rl.20

van Dinter, Marieke and Kim M. Cohen, Wim Z. Hoek, Esther Stouthamer, Esther Jansma, Hans Middelkoop, Late Holocene lowland fluvial archives and geoarchaeology: Utrecht’s case study of Rhine river abandonment under Roman and Medieval settlement, Quarternary Science Reviews, 166 (2017) 227- 265, http://dx.doi.org/10.1016/j.quascirev.2016.12.003

Pierik HJ. Landscape changes and human–landscape interaction during the first millennium AD in the Netherlands. Netherlands Journal of Geosciences, Volume 100, e11. https://doi.org/10.1017/njg.2021.8 , also https://www.academia.edu/51016582/Landscape_changes_and_human_landscape_interaction_during_the_first_millennium_AD_in_the_Netherlands?uc-g-sw=18993499

de Kleijn, M., Beijaard, F., Koomen, E., & van Lanen, R. (2018). Simulating past land use patterns: The impact of the Romans on the Lower-Rhine delta in the first century AD. Journal of Archaeological Science: Reports, 20, 244-256. https://doi.org/10.1016/j.jasrep.2018.04.006

[94] Gerlach R, Meurers-Balke J, Kalis AJ. The Lower Rhine (Germany) in Late Antiquity: a time of dissolving structures. Netherlands Journal of Geosciences. 2022;101:e14. doi:10.1017/njg.2022.11, https://www.cambridge.org/core/journals/netherlands-journal-of-geosciences/article/lower-rhine-germany-in-late-antiquity-a-time-of-dissolving-structures/4D37B48DA4EC00C60A83AF11AFDB7CFE, also https://www.academia.edu/108260441/The_Lower_Rhine_Germany_in_Late_Antiquity_a_time_of_dissolving_structures

Rowin J van Lanen and Bert Groenewoudt, Counting heads: Post-Roman population decline in the Rhine-Meuse delta (the Netherlands) and the need for more evidence-based reconstructions, in Niall Brady & Claudia Theune, eds, Settlement Change Across Medieval Europe: Old paradigms and New Vistas, , 2019, Ruralia XII, Leiden; Sidestone, 2019, https://www.academia.edu/40523376/Counting_heads_Post_Roman_population_decline_in_the_Rhine_Meuse_delta_the_Netherlands_and_the_need_for_more_evidence_based_reconstructions

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Altena, E., Smeding, R., van der Gaag, K.J. et al. The dutch Y-chromosome from the early middle ages to present day. Archaeol Anthropol Sci 17, 116 (2025). https://doi.org/10.1007/s12520-025-02224-4

[96] Iñigo Olalde, Pablo Carrión, Ilija Mikić, Nadin Rohland, Shop Mallick, Iosif Lazaridis, Miomir Korać, Snežana Golubović, Sofija Petković, Nataša Miladinović-Radmilović, Dragana Vulović, Kristin Stewardson, Ann Marie Lawson, Fatma Zalzala, Kim Callan, Željko Tomanović, Dušan Keckarević, Miodrag Grbić, Carles Lalueza-Fox, David Reich, Cosmopolitanism at the Roman Danubian Frontier, Slavic Migrations, and the Genomic Formation of Modern Balkan Peoples, bioRxiv 2021.08.30.458211; doi: https://doi.org/10.1101/2021.08.30.458211

Bird, Steven, Haplogroup E3B1A2 as a Possible Indicator of Settlement in Roman Britain by Soldiers of Balkan Origin, Journal of Genetic Genealogy, Volume 3, Number 2 (Fall 2007) Reference Number: 32.006, https://jogg.info/wp-content/uploads/2021/09/32.006.pdf

What Ancient DNA Reveals About the Medieval Population of the Low Countries, Medievalist.net, https://www.medievalists.net/2025/05/ancient-dna-low-countries/

Altena, E., Smeding, R., van der Gaag, K.J. et al. The dutch Y-chromosome from the early middle ages to present day. Archaeol Anthropol Sci 17, 116 (2025). https://doi.org/10.1007/s12520-025-02224-4

Olalde I, Altena E, Bourgeois Q, Fokkens H, Amkreutz L, Deguilloux MF, Fichera A, Flas D, Gandini F, Kegler JF, Kootker LM, Leijnse K, Kooijmans LL, Lauwerier R, Miller R, Molthof H, Noiret P, Raemaekers DCM, Rivollat M, Smits L, Stewart JR, Anscher TT, Toussaint M, Callan K, Cheronet O, Frost T, Iliev L, Mah M, Micco A, Oppenheimer J, Patterson I, Qiu L, Soos G, Workman JN, Edwards CJ, Lazaridis I, Mallick S, Patterson N, Rohland N, Richards MB, Pinhasi R, Haak W, Pala M, Reich D. Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups. bioRxiv [Preprint]. 2025 Mar 25:2025.03.24.644985. doi: 10.1101/2025.03.24.644985. PMID: 40196638; PMCID: PMC11974744 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11974744/

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Olalde I, Altena E, Bourgeois Q, Fokkens H, Amkreutz L, Deguilloux MF, Fichera A, Flas D, Gandini F, Kegler JF, Kootker LM, Leijnse K, Kooijmans LL, Lauwerier R, Miller R, Molthof H, Noiret P, Raemaekers DCM, Rivollat M, Smits L, Stewart JR, Anscher TT, Toussaint M, Callan K, Cheronet O, Frost T, Iliev L, Mah M, Micco A, Oppenheimer J, Patterson I, Qiu L, Soos G, Workman JN, Edwards CJ, Lazaridis I, Mallick S, Patterson N, Rohland N, Richards MB, Pinhasi R, Haak W, Pala M, Reich D. Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups. bioRxiv [Preprint]. 2025 Mar 25:2025.03.24.644985. doi: 10.1101/2025.03.24.644985. PMID: 40196638; PMCID: PMC11974744. (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11974744/

Altena, E., Smeding, R., van der Gaag, K.J. et al. The dutch Y-chromosome from the early middle ages to present day. Archaeol Anthropol Sci 17, 116 (2025). https://doi.org/10.1007/s12520-025-02224-4

Sasso, S et al.,, Capturing the fusion of two ancestries and kinship structures in Merovingian Flanders, Proc. Natl. Acad. Sci. U.S.A. 121 (27) e2406734121, https://doi.org/10.1073/pnas.2406734121 (2024).

Beneker, O., Molinaro, L., Guellil, M. et al. Urbanization and genetic homogenization in the medieval Low Countries revealed through a ten-century paleogenomic study of the city of Sint-Truiden. Genome Biol 26, 127 (2025). https://doi.org/10.1186/s13059-025-03580-z

Gretzinger, J., Sayer, D., Justeau, P. et al. The Anglo-Saxon migration and the formation of the early English gene pool. Nature 610, 112–119 (2022). https://doi.org/10.1038/s41586-022-05247-2

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Genetic History of Europe, Wikipedia, This page was last edited on 11 November 2025, https://en.wikipedia.org/wiki/Genetic_history_of_Europe

Altena, E., Smeding, R., van der Gaag, K.J. et al. The dutch Y-chromosome from the early middle ages to present day. Archaeol Anthropol Sci 17, 116 (2025). https://doi.org/10.1007/s12520-025-02224-4 see also https://d-nb.info/1371388660/34

[99] Olalde I, Carrión P, Mikić I, Rohland N, Mallick S, Lazaridis I, Mah M, Korać M, Golubović S, Petković S, Miladinović-Radmilović N, Vulović D, Alihodžić T, Ash A, Baeta M, Bartík J, Bedić Ž, Bilić M, Bonsall C, Bunčić M, Bužanić D, Carić M, Čataj L, Cvetko M, Drnić I, Dugonjić A, Đukić A, Đukić K, Farkaš Z, Jelínek P, Jovanovic M, Kaić I, Kalafatić H, Krmpotić M, Krznar S, Leleković T, M de Pancorbo M, Matijević V, Milošević Zakić B, Osterholtz AJ, Paige JM, Tresić Pavičić D, Premužić Z, Rajić Šikanjić P, Rapan Papeša A, Paraman L, Sanader M, Radovanović I, Roksandic M, Šefčáková A, Stefanović S, Teschler-Nicola M, Tončinić D, Zagorc B, Callan K, Candilio F, Cheronet O, Fernandes D, Kearns A, Lawson AM, Mandl K, Wagner A, Zalzala F, Zettl A, Tomanović Ž, Keckarević D, Novak M, Harper K, McCormick M, Pinhasi R, Grbić M, Lalueza-Fox C, Reich D. A genetic history of the Balkans from Roman frontier to Slavic migrations. Cell. 2023 Dec 7;186(25):5472-5485.e9. doi: 10.1016/j.cell.2023.10.018. PMID: 38065079; PMCID: PMC10752003. (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC10752003/

Iñigo Olalde, Pablo Carrión, Ilija Mikić, Nadin Rohland, Shop Mallick, Iosif Lazaridis, Miomir Korać, Snežana Golubović, Sofija Petković, Nataša Miladinović-Radmilović, Dragana Vulović, Kristin Stewardson, Ann Marie Lawson, Fatma Zalzala, Kim Callan, Željko Tomanović, Dušan Keckarević, Miodrag Grbić, Carles Lalueza-Fox, David Reich, Cosmopolitanism at the Roman Danubian Frontier, Slavic Migrations, and the Genomic Formation of Modern Balkan Peoples, bioRxiv 2021.08.30.458211; doi: https://doi.org/10.1101/2021.08.30.458211

Macjamo, Ancient DNA of Roman Danubian Frontier and Slavic Migrations (Olalde 2021), Iron Age & Antiquity Forum, Eupedia , https://www.eupedia.com/forum/threads/ancient-dna-of-roman-danubian-frontier-and-slavic-migrations-olalde-2021.41753/

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[100] Altena, E., Smeding, R., van der Gaag, K.J. et al. The dutch Y-chromosome from the early middle ages to present day. Archaeol Anthropol Sci 17, 116 (2025). https://doi.org/10.1007/s12520-025-02224-4 see also https://d-nb.info/1371388660/34

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[101] What Ancient DNA Reveals About the Medieval Population of the Low Countries, Medievalist.net, https://www.medievalists.net/2025/05/ancient-dna-low-countries/

Altena, E., Smeding, R., van der Gaag, K.J. et al. The dutch Y-chromosome from the early middle ages to present day. Archaeol Anthropol Sci 17, 116 (2025). https://doi.org/10.1007/s12520-025-02224-4

[102] Rui Martiniano, Bianca De Sanctis, Pille Hallast, Richard Durbin, Placing Ancient DNA Sequences into Reference Phylogenies, Molecular Biology and Evolution, Volume 39, Issue 2, February 2022, msac017, https://doi.org/10.1093/molbev/msac017

Rohrlach, A.B., Papac, L., Childebayeva, A. et al. Using Y-chromosome capture enrichment to resolve haplogroup H2 shows new evidence for a two-path Neolithic expansion to Western Europe. Sci Rep 11, 15005 (2021). https://doi.org/10.1038/s41598-021-94491-z

[103] Rui Martiniano, Bianca De Sanctis, Pille Hallast, Richard Durbin, Placing Ancient DNA Sequences into Reference Phylogenies, Molecular Biology and Evolution, Volume 39, Issue 2, February 2022, msac017, https://doi.org/10.1093/molbev/msac017

Rohrlach, A.B., Papac, L., Childebayeva, A. et al. Using Y-chromosome capture enrichment to resolve haplogroup H2 shows new evidence for a two-path Neolithic expansion to Western Europe. Sci Rep 11, 15005 (2021). https://doi.org/10.1038/s41598-021-94491-z

[104] What Ancient DNA Reveals About the Medieval Population of the Low Countries, Medievalist.net, https://www.medievalists.net/2025/05/ancient-dna-low-countries/

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[105] Similar challese exist in Britain: Scheib CL, Hui R, Rose AK, D’Atanasio E, Inskip SA, Dittmar J, Cessford C, Griffith SJ, Solnik A, Wiseman R, Neil B, Biers T, Harknett SJ, Sasso S, Biagini SA, Runfeldt G, Duhig C, Evans C, Metspalu M, Millett MJ, O’Connell TC, Robb JE, Kivisild T. Low Genetic Impact of the Roman Occupation of Britain in Rural Communities. Mol Biol Evol. 2024 Sep 4;41(9):msae168. doi: 10.1093/molbev/msae168. PMID: 39268685; PMCID: PMC11393495. (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11393495/

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