Looking at the Griff(is)(es)(ith) Y-DNA Phylogenetic Gap Associated with the Meuse and Rhine River Watershed from the Bronze Age Onward – Part Five

This story focuses on the examination of possible ecological, demographic, and social-cultural influences that may explain the lack of identified subclades (YDNA ancestors) in the migratory path of the Griffis genetic paternal line that lived in the Meuse Rhine watershed area. The story discusses possible social and cultural groups that may have been associated with the generations of ancestors of the Griff(is)(es)(ith) that may have lived during and after the bronze age, up to approximately 500 BCE.

The 2,850 year Gap between G-FGC7516 and G-Z6748: The most common recent ancestor associated with G-FGC7516 was born around 2200 BCE. The next genetic ancestor on the Griff(is)(es)(ith) YDNA line was associated with the genetic SNP mutation defining the G-Z6748 haplogroup, 2,850 years later. This gap of undocumented YDNA ancestors represents about 95 generations.

The absence of identified subclades between haplogroups G-FGC716 (c. 2200 BCE) and G-Z6748 (c. 650 CE) in the Meuse-Rhine watershed area are likely the results from interconnected social, demographic, and environmental factors and their successive effects over generations . Some of the factors that may have influenced the lack of identified haplogroups along this migratory path are:

  • Environmenal factors as well as targeted and genetic sampling strategies; .
  • The persistant interaction between social and cultural groups within the watershed area; and
  • The long term effects of social and cultural diversity in the watershed area.

Environmental and Methodological Factors

The retrieval of ancient DNA (aDNA) in the Meuse Rhine watershed area is highly contingent on environmental factors such as flooding and sediment dynamics, soil chemistry, climate cycles, and human land use, as well as targeted archaeological and genetic sampling strategies that mitigate adverse preservation conditions. Archaeological sites that are well preserved will yield promising evidence to reconstruct ancient social and cultural practices as depicted in illustration one.


Illustration One: Example of an Archaeological Site and aDNA Retrieval and Analysis

Click for Larger View | Source: Oğuzhan Parasayan et al. , Late Neolithic collective burial reveals admixture dynamics during the third millennium BCE and the shaping of the European genome. Sci. Adv . 10, eadl2468 (2024). DOI:10.1126/sciadv.adl2468, https://www.science.org/doi/10.1126/sciadv.adl2468

(A) Map showing geographical locations of samples. (B) Estimated age of death and haplogroup information of the individuals. (C) Pedigree plots showing kin relationships between the individuals. (D) Pedigree plots showing the genomic component of steppe-related and Late Neolithic ancestries.


The Meuse and Rhine delta regions, acting both as a conservator and genetic crossroad, provides a broader spectrum of recoverable lineages, including indigenous and incoming populations. The upland and interior Rhine and Meuse sites are more prone to genetic bottlenecks and loss due to environmental barriers and patchy skeletal survival.

Illustration Two: Migratory Path and the Meuse and Rhine Rivers

Click for Larger View | Source: Modified version of map found in Pilarczyk, Krystian. (2007). NATO Science Series. 10.1007/978-1-4020-5741-0_26. ,Fig. 1 The Rhine and Meuse basin Page 26 ,https://www.researchgate.net/figure/The-Rhine-and-Meuse-basin_fig11_226598873

Illustration Three: The Meuse and Rhine Watersheds

Click for Larger View | Source: Verkade, Van Simon, J.D. Brown, Femke Davids, Estimating predictive hydrological uncertainty by dressing deterministic and ensemble forecasts; a comparison, with application to Meuse and Rhine, Oct 2017, Hournal of Hydrology, 555:257-277, DOI: 10.1016/j.hydrol.2017.10.024

The region’s humid and waterlogged conditions, driven by fluctuating river regimes and frequent flooding, generally accelerate the degradation of skeletal remains and ancient DNA (aDNA) through microbial activity, hydrolysis, and oxidation. This results in poor long-term preservation. These processes could either bury remains in anoxic, sediment-rich environments conducive to preservation or expose them to further decay. Upland settlement patterns influenced DNA retrieval potential. Archaeological sites located in upland sandy soils were more likely to yield degraded samples due to acidity while delta wetlands offered better DNA survival prospects due to waterlogging and reduced oxygen. [1]

Human environmenal activities through this particular historical period had impacts on the preservation of aDNA. Major land use changes, such as deforestation, agricultural practices, the historical layering of land use practices, and Roman-medieval infrastructure changes (e.g. ditches, embankments), altered erosion rates and sedimentation. This sometimes created sealed burial contexts that were beneficial for DNA preservation but could also disturb existing archaeological layers and lead to destruction or mixing of deposits. Increased flooding events, especially from the Iron Age onward and during the Roman and early Medieval periods, redistributed sediments, alternately exposing and sealing archaeological contexts. [2]

Table One: Environmenal Factors Affecting DNA Retrieval in the Meuse Rhine Watershed Area

FactorEffect on DNA
Humidity & WaterloggingAccelerates decay
Flooding & Sediment DynamicsBurial or exposure
Soil pH (Alkaline vs. Acidic)Preserves vs. degrades DNA
Climate OscillationsCooling preserves, warming degrades
Land Use Changes (Deforestation, Ditches)Mixed impact

The Rhine-Meuse delta’s flood-prone geography disrupted settlement continuity. Repeated flooding could isolate populations, causing local extinctions of lineages. Prolonged reliance on localized wetland resources limited population mobility and genetic exchange.

The Barbed Wire Beaker Culture and the Archaeological Concept of Cultural Change (~ 2100 – 1800 BCE)

The beginning of this phylogenetic gap of YDNA haplogroups in the Griff(is)(es)(ith YDNA line of descent begins with an individual who had a 68 percent chance of being born between 2533 BCE and 1853 BCE. Based on inferences from YDNA mapping data [3] , I have suggested that this most common recent ancestor of haplogroup G-FGC7516 lived in a general area that is now Aachen, Germany.  Aachen is located west of the Rhine river and is part of the Meuse-Rhine watershed area. (see illustration one). The city sits on the Wurm River, a tributary of the Rur River, which flows into the Meuse river system. [4]

Illustration One: Location of Aachen Germany

Click for Larger View

Our most recent common ancestor could have lived in a community that, in contemporary times, would be labeled by achaeologists as part of the Barb Wire Beaker or Bell Beaker culture.

The Barbed Wire Beaker culture was a local development of the Bell Beaker phenomenon in the Meuse-Rhine watershed area, specifically around 2100–1800 BCE. The Barbed Wire Beaker culture arose locally from the Bell Beaker culture and is archaeologically documented specifically in the lower Rhine and Meuse river areas, which today comprise the southern Netherlands and adjacent western Germany. [5]

This time period in the Meuse Rhine watershed area was a period of cultural transition. Social and cultural characteristics of the Barbed Wire Beaker included a distinctive pottery style, continued pastoralist and agricultural traditions, and the use of burial mounds, reflecting both continuity with Late Neolithic practices and early Bronze Age innovations. The continued use of burial mounds (barrows) for the interment of individuals was a key feature, with evidence for both inhumation and, to a lesser extent, cremation, hinting at social differentiation based on grave goods and monument size. [6]

As in neighbouring countries, the period between 2000 and 1800 BC shows continuity with the Late Neolithic Beaker Cultures. . . . Bell Beaker forms and decorative patterns continued in Barbed Wire Beaker pottery . . . , even in the characteristic large beaker pots. The only difference with Bell Beakers appears to be the decrative technique. Instead of using a spatula, barbed wire decoration is made using a flexible or sturdy stamp with a piece of string wrapped around it which is pressed into the wet clay. The distribution of Barbed Wire pottery also resembles that of late Bell Beaker closely to the extent that it is not found in the higher parts of Belgium and Germany. It extends into the higher regions only along the river valleys of the Rhine and the Meuse. [7]

Harry Fokkens critically analyzes long-standing assumptions about the origins of the Bell Beaker phenomenon in the Netherlands. He challenges the typological and chronological sequence that underpins the so-called ‘Dutch Model’ in archaeology. The Dutch Model argues for a local, continuous development from Single Grave Beakers culture into Bell Beakers culture, positioning the Rhine-Meuse delta as a key formative region. Fokkens scrutinizes the reliance on pottery typology and 14C carbon dating as primary evidence for cultural continuity. He argues that these are insufficient markers for true cultural identity and for explaining the dynamics of change.

He points out that settlement data do not fully support the Dutch Model. There are signs of regional cultural diversity and discontinuity, especially when considering earlier traditions like the Vlaardingen and Funnel Beaker cultures. He calls for moving beyond typological sequence and grave goods, advocating for the study of how beaker practices were incorporated into existing local traditions, a shift toward exploring the diffusion of innovations rather than migration as the sole explanatory frame. [8]

Illustration Two: Model of Adoption of Innovations [9]

Click for Larger View

Color intensity indicates archaeological visibility of objects  & features. (A) Visibility of subsequent cultural traditions; (B) Shows how introduction trajectory of innovations may cause differential visibility & subsequent archaeological perception of abrupt change.

Harry Fokkens defines cultural change not simply as a typological sequence in material culture or as abrupt migration, but as a process that must be interpreted through a wider lens—including settlement data, local background, and context-driven adoption of practices. He emphasizes moving beyond static models, arguing that culture change requires understanding the social and historical processes that drive adoption and transformation—settlement archaeology, agency, and local adaptation are key (see illustration two). Cultural change should be studied by examining how new practices fit into everyday life, ritual, and economy, not just as visible shifts in craft or burial customs.

Cultural change, for Fokkens, is defined by how innovations (such as Bell Beakers) are adopted, adapted, and integrated into local traditions across different regional contexts. This can include diffusion, selective adoption, or hybridization, making each region’s trajectory unique.

Major change only develops when the critical mass is reached and the rate of adoption changes (see illustration two) . . . . This is the point where so many people have already adopted an innovation that non-adopters run the risk not to belong to the ‘mainstream’ any longer. . . . I have suggested that this is the phase that innovations become archaeologically visible and that we may ‘see’ culture change. In my view this transition phase, the period of adoption before the critical mass is reached, can be recognised in many regions of Europe. This is characterised as a period in which new (Bell Beaker) elements are incorporated in regional traditions without changing them. . . . So regionalisation may well have started 2400 cal BC or earlier, but had developed around 2300 cal BC.” [10]


The map in illustration three visually depicts the cultural landscape of the Netherlands between 2500-2000 BCE as a period of transition. The map depicts different variants of the Bell Beaker cultures that may have existed simultaneously and overlapped over time.

A period that our common YDNA ancestor and susequent generations could have existed. In illustration three, known, discovered Bell Beaker sites are presented along with other cultural sites. The map depicts the distribution of barrow grave sites, Veluvian Bell Beaker sites and Notheast Dutch/German Beaker sites. The map also depicts the distribution of maritime Bell Beaker type archaelogical sites.

Illustration Three: The cultural landscape of the Netherlands c. 2500-2000

Click for Larger View | Source: Figure 9 in Harry Fokkens, Background to Dutch Beakers A Critical Review in  Fokkens, Harry & Franco Nicolis, (eds), Background to Beakers Inquiries into Regional Cultural Backgrounds of the Bell Beaker Complex, Leiden: Sidestone Press, 2012, 9-36, https://www.sidestone.com/openaccess/9789088900846.pdf

Fokkens contends that determining the characteristics of archaeological culture and cultural change are better understood through the exploration of local settlement contexts rather than just burial typology or other singular changes in artifacts. He encourages a multidisciplinary approach: looking at settlement archaeology, regional interactions, and innovations in local contexts to reinterpret the rise and spread of Bell Beaker culture in the Netherlands.

Illustration four provides two maps of Bell Beaker and Barbed Wire cultural sites and burial grounds. The comparison of the two maps highlight the overlap of the two archaeological cultures and their location on waterways in the Meuse Rhine watershed.

Illustration Four: Bell Beaker and Barb Wire Culture Sites

Click for Larger view | Source: Pat of Figure 3.13 in Fokens, H. , B.J.W. Steffens & S.F.M. van As, Farmers, fishers, fowlers, hunters Knowledge generated by development-led archaeology about the Late Neothithic, the Early Bronze Age and the start of the Middle Bronze Age (2850 – 1500 cal BC) in the Netherlands, Amersfoort: Rijksdienst voor het Cultureel Erfgoed, 2016, Page 38,https://collectie.huisvanhilde.nl/pdf/rce_nar-053_farmers_fishers_fowlers_hunters.pdf

Community and family units in the Dutch Beaker cultures appear to have been based in small, dispersed settlements with some degree of sedentism, engaging in mixed farming, pastoralism, and hunting as indicated by archaeological settlement finds and faunal remains. Domestic sites show continuity in housing forms from the Late Neolithic, with initial persistence of two-aisled houses before a transition to three-aisled “longhouse” structures in the following centuries. [11]

Throughout the 20th century, scholars debated whether the Beaker phenomenon in northwestern Europe was best explained by waves of migrating peoples (possibly a small elite or specialists) or by the widespread adoption of ideas and artifacts across existing societies. The migrationist perspective dominated early on, but after the 1960s, archaeologists increasingly favored diffusionist models, seeing migration as a last resort explanation. Renewed attention to migration has emerged in recent decades with the advent of aDNA studies, which have provided new insights into population movements. [12]

Migrationists argued that significant cultural transformations, like the appearance of the Beaker culture’s characteristic pottery and burial customs, reflected the arrival or influx of new populations. In the Beaker context, this could mean the movement of groups such as warriors, craftsmen, or traders, who settled in new regions and brought their traditions, technologies, and genetic lineages with them.

Diffusionists emphasized that similar artifacts or cultural traits could spread via contact, trade, or imitation without mass population movement. For the Beaker culture, this meant that local populations could adopt Beaker pottery styles, burial rites, or metalworking techniques through networks of exchange and social influence, rather than as a result of being replaced or displaced by newcomers.

Recent studies combining archaeology and ancient DNA analysis have led to nuanced conclusions regarding the migrationism versus diffusionism debate about the Beaker culture in northwestern Europe. The general consensus is that both migration and diffusion played significant but contextually distinct roles. [13]

The Bell Beaker-associated populations in the Meuse–Rhine region (including the Netherlands, Belgium, and western Germany) emerged through a mixture of local inhabitants and Corded Ware-associated migrants. As indicated in previous parts of this story, the Meuse–Rhine region stands out due to its long persistence of hunter-gatherer and early European farmer genetic ancestry—up to three millennia longer than neighboring continental areas, as wetlands and riverine ecology limited gene flow and farming adoption. The first appearance of Bell Beakers in this region marked a rapid shift in genetic makeup and cultural practices, distinct from earlier local continuity. [14]

The Barbed-Wire Beaker culture marks the beginning of the Bronze Age in the Dutch river area. Metalworking (especially the introduction of bronze items) is evident but limited, suggesting access via long-distance exchange rather than local production. Prestige items (e.g. axes, daggers) entered the region as exotic goods. The Netherlands lacked the natural copper and tin resources needed to produce bronze. Consequently, all bronze objects were either imported or created from imported alloys, demonstrating the region’s connection to larger European trade networks. This timing aligns with the initiation of Bronze Age settlement and metallurgy in the broader Dutch river area, including both the Meuse and the lower Rhine valleys. [15]

Bell Beaker people took advantage of transport by sea and rivers, creating a cultural spread extending from Ireland to the Carpathian Basin and south along the Atlantic coast and along the Rhône valley to Portugal, North Africa, and Sicily, even penetrating northern and central Italy. [16]

Barbed Wire Beaker communities participated in extensive riverine trade and communication networks, which would almost certainly have required effective watercraft such as logboats. The spread of beaker styles and goods along major waterways (Rhine, Meuse, IJssel) is consistent with an environment where logboats were in routine use, facilitating cultural exchange. [17]

. . . (I)nstead of with ‘cultures’, we are dealing with different communities of practice that through regular communication were structured around learned practices. This notion shifts the focus of study from objects to communication networks and different way of ‘doing things’. Many of the sites excavated the last 15 years and before were located in dynamic landscapes and near rivers or lakes, even near the sea . Therefore we expect water ways to have structured the cultural palette and the transmission of ideas. From that perspective we predict that river deltas and upstream their catchment areas constituted the natural routes for transport of objects and people and the foremost connections in the communication networks. Land-based connections should of course not be ignored, but they may not have been of primary importance for structuring and maintaining local communities. [18]

“. . . (T)he location of many settlement in the dynamic landscapes and on the borders of wetlands may also imply that transport and travel by boats and canoes was far more important than with land-based vehicles. Wheels and carts were known, but long-distance communication over land seems unviable because these burdens were probably too heavy We suggest that instead waterways were by far the most important means of transport and communication. This directly influences our ideas about the constitution of regional groups as communities of practice. [19] (Emphasis is mine.)

Logboats, also known as dugout canoes or monoxylons, are simple watercraft made by hollowing out the trunk of a single tree. They are among the oldest known types of boats, with archaeological examples dating back over 8,000 years to the Neolithic era. [20] Logboats represent a key technological innovation in early water transport. The Meuse-Rhine system formed a natural corridor for movement and exchange stretching from the interior of Europe to the North Sea, and logboats provided the practical technology to exploit these routes.

Archaeological evidence suggests that some of the earliest regional trade networks, including the distribution of pottery styles, flint, and metal objects, relied on water transport made possible by logboats. Logboats were essential for fishing, hunting, and gathering in wetland zones, as well as for accessing and transporting timber, peat, and other bulk resources typically found along watercourses. Radiocarbon-dated finds show that logboats were used continuously from the Mesolithic into the Bronze Age and later periods in the Meuse-Rhine watershed, highlighting their enduring relevance. The region’s dynamic fluvial landscape—with frequent avulsions, flooding, and marsh formation—made watercraft indispensable for everyday life. [21]

“The Pesse canoe is the world’s oldest – known boat. Carbon dating indicates that the boat was constructed during the early Mesolithic period between 8040 BC and 7510 BC.” [22]

Illustration Five: Pesse Logboat

Click for Larger View | Source: Christoph Braun, Boomstamkano van Pesse, 12 April 2016, Wikimedia Commons, https://commons.wikimedia.org/wiki/File:Boomstamkano_van_Pesse,_Drents_Museum,_1955-VIII-2.jpg

Settlement clusters frequently appear along major rivers and tributaries, with evidence that logboats enabled frequent movement and social ties among dispersed communities. Some prehistoric villages in the Rhine-Meuse delta are interpreted as trading posts or hubs precisely because of their strategic waterfront locations and the availability of logboat transport.

An interesting study by Lanting presents the largest radiocarbon and dendrochronological dataset for European logboats. It covers more than 600 directly dated specimens across Europe with additional meta-analysis of over 3,500 archaeological finds of logboats. The study establishes a clear chronological and geographic set of diffusion patterns associated with one form of water transportation. (see illustration six). [23]

Illustration Six: Diffusion of Log Boats

Click for Larger View | Source: Map of Diffusion of Logboats,Lanting, J.N. 1997/98, Dates of origin and diffusion of the European logboat, Palaehistoria 39/40, P 640, https://www.ancientportsantiques.com/wp-content/uploads/Documents/ETUDESarchivees/Navires/Documents/Lanting2000-Logboats.pdf

When working with the dates of the logboats . . . some patterns in distribution both in space and time are noticeable. To visualize these patterns Continental Europe north of the Alps and the Pyrenees, and west of the Russian border has been divided in two zones . . . . Zone l comprises Denmark, northwestern Germany (i.e. Schleswig-Holstein, Hamburg, Niedersachsen, Bremen and Nordrhein-Westfalen), Netherlands, Belgium and northwestern France (i.e. the regions Nord-Pas-de-Calais, Picardie, Hauteand Basse-Normandie, Ile-de-France, ChampagneArdenne and the departments Eure-et-Loir and Meuse). Zone 2 comprises the rest of France and Germany, Switzerland, Austria, the Czech Republic and Poland.[24]

Robert Van de Noort’s article, “Exploring Agency Behind the Beaker Phenomenon”, emphasizes the central role of individual agents, particularly navigators, in creating and maintaining the long-distance exchange networks that underpinned the spread of the Bell Beaker phenomenon across Europe and North Africa. Instead of focusing on migration or simple diffusion models, Van de Noort highlights the active social practices and ritualized journeys of ‘elite travelers’ as key drivers of Beaker cultural expansion and transformation. Van de Noort draws on Mary Helms’s anthropological work, proposing that travel was a significant empowering activity giving travelers esoteric knowledge and access to exotic goods, which in turn conferred social power. His work reframes the Beaker phenomenon as a dynamic process shaped by individual agency, social networks, and long-distance travel practices, rather than mere migration or material diffusion. [25]

There is currently no evidence that logboats were a major feature or symbolic object of the Barbed Wire Beaker culture. They appear in the archaeological record as practical watercraft rather than cultural markers. The distribution of Barbed Wire Beaker settlements shows a preference for river valleys, lakes, and wetlands, possibly implying the routine use of logboats for transport, fishing, and access to dispersed settlements. Organic objects such as logboats rarely survive in grave or habitation contexts attributed explicitly to Barbed Wire Beaker sites, so direct association is difficult to prove archaeologically. [26]

The Elp and Hilversum Cultures (~ 1870 – 650 BCE)

In the Bronze Age, several regional cultures existed within the Meuse-Rhine watershed area, generally falling under broader classifications like the Elp culture and the Hilversum culture, in what is now the Netherlands and neighboring regions. These cultures inherited and further developed material traditions and social practices from the late Bell Beaker period, marking the beginning of the regional Bronze Age (see illustration seven).

Illustration Seven: Map of the Hilversum and Elp Cultures [27]

The Elp Culture was located primarily in the Netherlands and Northwestern Germany. It existed during the Middle and Late Bronze Age, roughly 1800-800 BCE. It is characterized by several distinctive archaeological features, especially in pottery, burial customs, settlement architecture, and subsistence strategies. The Elp culture is identified by its low-quality earthenware pottery known as “Kümmerkeramik” or “Grobkeramik,” which is considered an archaeological marker of the culture. This pottery style reflects an utilitarian approach, with little decorative emphasis. [28]

Elp settlements focused heavily on cattle raising, particularly for milk and milk products. Archaeological evidence suggests social importance and possibly ritual aspects tied to cattle, such as stacks of cowhides in graves and animal offerings. The stalling of cattle also points to adaptations to climatic conditions affecting milk production, as well as a potential response to social pressures like cattle raiding. [29]

Geographically, the Elp culture was part of the Nordwestblock, situated north and east of the Rhine and IJssel rivers and bordered by the Hilversum and Hoogkarspel cultures. These cultures, including Elp, are believed to have derived from the earlier Bell Beaker culture, thus placing the Elp culture at the cultural and ecological interface between North Sea (Atlantic) and Northern (Nordic) horizons. [30]

Based on archaeological study, the Elp culture is distinguished within the context of Middle and Late Bronze Age northwestern Europe. The early phase (1800–1200 BCE) is characterized by Tumuli burials, two to three-aisled longhouses (see illustrations eight and nine), and strong links to Tumulus culture. [31] The late phase (1200–800 BCE) is characterized by the transition to Urnfield culture, increased collective cremations, material and settlement continuity into the Early Iron Age. [32]


Illustration Eight: Two and Three Aisle House Construction in the Meuse Rhine Watershed Area

Longhouses were typically multifunctional, serving as both living quarters for humans and stalls for livestock, often with a central space for work. [33]

Structural elements of a house and their names A: a two-aisled construction; B: a three-aisled construction

Click for Larger View | Source: Figure 6.3 in Fokens, H. , B.J.W. Steffens & S.F.M. van As, Farmers, fishers, fowlers, hunters Knowledge generated by development-led archaeology about the Late Neothithic, the Early Bronze Age and the start of the Middle Bronze Age (2850 – 1500 cal BC) in the Netherlands, Amersfoort: Rijksdienst voor het Cultureel Erfgoed, 2016, Page 38 , https://collectie.huisvanhilde.nl/pdf/rce_nar-053_farmers_fishers_fowlers_hunters.pdf

Illustration Nine: Reconstruction drawing of a Late Bronze Age Longhouse

Click for Larger View | Source: Kristian Kristiansen, Kristian. (2010). Decentralized Complexity: The Case of Bronze Age Northern Europe. 10.1007/978-1-4419-6300-0_7. Fig. 7.12 Reconstruction drawing of a Late Bronze Age hamlet from Holland, Pages 169-192, in T Douglas Price and Gary M. Feinman, eds, Pathways to Power: New Perspectives on the Emergence of Social Inequality, New York: Springer, 2010;https://www.researchgate.net/publication/225207647_Decentralized_Complexity_The_Case_of_Bronze_Age_Northern_Europe

The Hilversum Culture dominated the southern region of the Netherlands during the Middle Bronze Age (1800-800 BCE). Centered in the southern Netherlands and parts of northern Belgium, with archaeological influence extending into northern France, it also appears to have maintained cultural ties with the Wessex culture in Britain.  [34]

Current studies suggest that the Hilversum Culture was characterized by extended family (possibly patrilocal and patrilineal) households centered in large longhouses, with evidence of kinship-based organization typical of northwest European Bronze Age societies. The social structure appears to have revolved around extended kin groups occupying substantial farmsteads, combining people and livestock under a single roof, which provided both economic and social cohesion. [35]

Archaeological evidence indicates a trend toward large and even very large farmhouses, likely hosting extended families (kin groups spanning multiple generations). Although direct ancient DNA from the Hilversum region is limited, regional analogs and house organization suggest married daughters often left their natal homes to join their husband’s family group, supporting a patrilocal system; inheritance and lineage may have been patrilineal, as seen elsewhere in contemporary northwestern Europe. [36]

The centrality of kinship and the presence of large houses imply that rights to land, resources, and the leadership of family units were inherited, probably following male lines. The inclusion of cattle within the domestic sphere, both economically and ideologically, reinforced the importance of kin-based labor and resource management, further binding family units together. [37]

Patrilineal and patrilocal practices may have had a continued negative impact on subclade proliferation on minority lineages such as the Griff(is)(es)(ith) line. Based on the time period associated with the Hilversum culture and the estimated migratory path (see ilustration ten), it is likely that undocumented generations of the YDNA lineage of the Griff(is)(es)(ith) paternal line were part of this culture.

Illustration Ten: The Netherlands as a Border Zone Belween the Atlantic and Nordic Worlds – Griff(is)(es)(ith) paternal YDNA migratory path ~1850 – 1650 BCE

Click for Larger View | Source: Modification of a map originally inFokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001, uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

The distribution of the cultural differences in the Bronze Age shows that the boundary between the Nordic and the Atlantic worlds runs slightly erratically through the Netherlands (illustration ten above). It does not follow the courses of the Meuse and the Rhine when they start to flow westward near Nijmegen, but the river IJssel, a northern branch of the Rhine. So in many respects the eastern Netherlands are part of the Nordic world, whereas the Veluwe – the ice-pushed hilly area north of the Rhine and west of the IJssel – is still part of the Atlantic world. On the other hand, the coastal zone north of the Rhine and Meuse, including West Friesland, seems to be part of the Nordic world in some respects . . . and of the Atlantic world in others, e.g. pottery . . . . Here probably navigation form the north along the coast played a role in the maintenance of contacts.[38]

The Tumulus Culture (~ 1600 – 1200 BCE)

Based on the migratoy path of the roughly 94 undocumented generations of the Griff(is)(es)(ith) YDNA line, some of the generations may have been part of the Tumulus culture. The Tumulus culture was the dominant material culture in Central Europe during the Middle Bronze Age, around 1600 to 1300 BCE. It was the descendant of the Unetice culture. Its heartland was the area previously occupied by the Unetice culture, and its territory included parts of Germany, the Czech Republic, Austria, Switzerland, the Carpathian Basin, Poland and France. [39]

The Tumulus culture was present in the eastern parts of the Meuse-Rhine watershed area during the Middle Bronze Age, though it was not its core territory (see illustration eleven). The Tumulus culture spread from its origins in central Europe to include the upper and middle Rhine valleys. The culture’s main heartland was farther east, centered in southern Germany, and expanded westwards, reaching as far as Alsace, France. 

Illustration Eleven: The Tumulus Culture and the Phylogenetic Gap

Click for Larger View | Source: Xoil, Europe during the Middle Bronze Age (simplified), 29 Jun 2024, Wikimedia Commons ,https://commons.wikimedia.org/wiki/File:Tumulus1a.png

Excavations have confirmed the existence of Tumulus culture sites in areas like the Ardennes Crest, which separates the Meuse and Rhine drainage basins. For example, one 1969 expedition worked on a tumulus (burial mound) at Ebly, located just within the Rhine’s drainage area. [40]

The Tumulus culture was fundamentally a warrior society, with social hierarchies centered on powerful chiefdoms and elite groups. These elites displayed status through weapon grave goods (such as swords, spears, and daggers) and the construction of monumental burial mounds, indicating the rise of a military aristocracy and decentralized kin-based groups. [41]

Illustration Twelve: Rendition of Individual from Tumulus Culture

Click for Larger View | Source: Modified image of Daderot, Gentleman, Bronze Age, 15th century BC, replica – Naturhistorisches Museum Nürnberg – Nuremberg, Germany,24 Oct 2016, Wikimedia Commons, https://commons.wikimedia.org/wiki/File:Gentleman,_Bronze_Age,_15th_century_BC,_replica_-_Naturhistorisches_Museum_Nürnberg_-_Nuremberg,_Germany_-DSC04215.jpg

The Tumulus culture helped shape a pan-European martial tradition, stimulated technological innovation in weapons, and made warfare a central feature of social and political life through the institutionalization of a warrior elite and advances in fortification architecture. Armament burial and iconography (such as grave steles and rock carvings) suggest that warfare became an integral part of male identity and community structure. Archaeological evidence, including skeletal trauma, weapons embedded in remains, and mass grave sites, points to an increase in violence and the normalization of warfare in social life. [42]

In the Meuse-Rhine area, archaeological evidence exists that lends credence to some forms of social stratification conveyed through artifacts of funerary rituals and burial mounds, with community constructed mounds as territorial markers and symbols of power. [43]

Tumulus culture communities were involved in extensive trade across central Europe, exporting items like amber and bronze goods and importing good from the Mediterranean area. Social status and identity were archaeologically reflected not only by individual grave goods but also by the organization of settlements and ritual landscapes, such as fortified hilltops or large necropoleis. Symbolism related to solar cults and ancestor veneration is inferred from grave architecture, metal objects, and ritual practices, suggesting ideological innovation and the transmission of new cultic beliefs. [44]

Based on acrchaaelogical studies, the Tumulus culture had an influence in the Meuse-Rhine watershed—especially in burial customs and elite symbolism—mainly through cultural diffusion along river routes rather than full-scale migration. This influence is documented in specialized archaeological and radiocarbon dating studies focused on Bronze Age barrow graves in the region. [45]

Based on archaeological evidence, the influence of Tumulus culture influence was likely less pronounced in the Meuse-Rhine delta than in the culture’s core regions (e.g., southern Germany and Bohemia), resulting in a mixture of local and intrusive burial and social patterns. [46] Overviews such as in the Oxford Handbook of the European Bronze Age describe a “northern branch” of the Tumulus phenomenon, which encompassed the lower Rhine and possibly parts of the Meuse watershed. [47] General comments can also be found regarding pan-European processes that highlight the Tumulus culture’s role in propagating warrior elite symbolism and funerary customs that reshaped the societies along the North-West European river systems during the Bronze Age. [48]

Archaeological evidence from Belgium and the southern Netherlands document the spread of Tumulus burial mounds (tumuli) during the Middle Bronze Age into areas bordering the Meuse-Rhine watershed, especially the eastern and southern edges of the Nordwestblock and adjacent Scheldt basin. [49]

Some of the undocumented generations and subclades of the Griff(is)(es)(ith) paternal line may have been part of this culture. Since the Tumulus culture was patrilineal and patrilocal and warlike, it is likely that the diversity of YDNA subclades was further reduced, creating a clear genetic signature of the kinship groupings and the reducing of subclades from minority groups. The concentration of men within their natal lineages and communities may have lead to a decrease in the overall variety of Y-chromosomes within those groups. 

While genomic studies have analyzed remains from Tumulus culture in Central Europe, such as southern Germany and northeastern France, at the time of writing this story, the studies do not include sampled burials found directly from the Lower Rhine, Netherlands, or adjacent Western German regions within the Meuse–Rhine drainage. Existing overviews reference the Tumulus culture’s archaeological expansion into northwest Europe, but ancient Y-DNA results from these westernmost areas remain absent in current literature. [50]

The Broader Urnfield Culture and the Lower Rhine Urnfield Culture (~ 1300 – 750 BCE)

The Tumulus culture in this region, as in much of Central Europe, was succeeded by the Late Bronze Age Urnfield culture, which also encompassed the area. The broader Urnfield culture continued to expand westward, bringing Proto-Celtic elements to the area. [51] Recent genomic studies propose that populations associated with Central European Urnfield culture contributed ancestry to later proto-Celtic groups across Europe, though the mechanism may have been centuries of gradual admixture rather than sudden mass movement. [52]

Illustration Thirteen: The Urnfield Culture Around 1300 BCE

Click for Larger View | Source: Simeon Netchev, Map of the Urnfield Culture c. 1300 BCE 23 August 2021, World History Encyclopedia, https://www.worldhistory.org/image/14456/map-of-the-urnfield-culture-c-1300-bce/

Generations of the the Griff(is)(es)(ith) paternal YDNA line may have been associated with the Lower Rhine Urnfield culture. The Urnfield Culture, while not strictly a regional culture specific to the Meuse-Rhine area, did spread throughout central and western Europe during the Late Bronze Age and influenced customs in the region, particularly in the southern Netherlands, see illustration ten.

“(U)rnfelds in the southwestern part of the Lower‑Rhine‑Basin are believed to fall under different cultural spheres of infuence: The so‑called ‘Atlantic Group’ orientated on the Belgian, French and English coast . . . and the ‘groupe Rhin-Suisse-France orientale’ that shows some cultural infuences of Central European urnfeld groups . . . . Clearly, the Lower‑ Rhine‑Basin at the time of the urnfelds is considered to have been a patchwork of small cultural entities (see illustration fourteen). “[53]

Illustration Fourteen: A Patchwork of ‘Small Cultural Entities’ Associated with the Urnfield Culture

Click for Larger View | Source: Arjan Louwen, Breaking and Making the Ancestors: Piecing together the urnfield mortuary process in the Lower-Rhine-Basin, ca. 1300 – 400 BC, Leiden, Slidestone Press, 2021, Page 13

Illustration Eleven shows the approximate migratory path of the Griff(is)(es)ith) paternal line in context of the distribution areas of the so-called ‘Ems-Group’ (North) and the ‘Niederrheinische Grabhügel Kultur’ (South) and their division into subgroups according to Verlinde and Hulst (2010). The black arrows represent the ‘cultural spread’ of the urnfelds from the presumed core areas near Münster and Rhineland in modern Germany. 1.) ‘Brabant-group’; 2.) ‘Niers-group’; 3.) ‘Veluwe-Utrecht-Gooi-group’; 4.) ‘Achterhoek-group’; 5.) ‘North-Netherlands- group’; 6z+n.) ‘Ems-group’ North (n) and South (z).

The Lower Rhine Urnfield culture (LRU) was a local variety of the broader Urnfield tradition, prominent in the Lower Rhine and Dutch river Delta regions during the Late Bronze Age (c. 1300–750 BCE). This culture is known for, among other things, its characteristic practice of cremating the dead and placing their ashes in pots (urns), which were then buried collectively in large fields—hence the term “Urnfield”.

The Lower Rhine Urnfield culture originated along the Rhine River and its delta, covering present-day regions in western Germany and the eastern Netherlands. Unlike other local groups, its cemeteries continued into the Early Iron Age.

In most parts of continental Europe, the first appearance of urnfields marks the beginning of a new archaeological period: the Late Bronze Age. The development of large cemeteries, often with hundreds of cremation graves, signified a fundamental break with the burial practice of the earlier period: a single inhumation or cremation grave covered by an earthen burial mound. At the same time many new types of pottery were introduced which in fabric, form and decoration differed completely from their Middle Bronze Age predecessors.[54]

Illustration Fifteen: Features of Urnfield Burials

Click for Larger View | Schematic overview of the types of archaeological features associated with urnfield graves and their terminology [55]

Urnfield-type burials refers to a Bronze Age burial practice, primarily in Central Europe, where the dead were cremated and their ashes were placed into ceramic urns, which were then interred in flat cemeteries. Named after the discovery of such urns in field-like burial grounds, this practice was characteristic of the Urnfield Culture and involved placing personal items or offerings with the cremains. The practice signifies a shift in beliefs, with cremation suggesting the physical form of the body was less relevant for the afterlife. This practice was a hallmark of the Urnfield culture, which emerged in the late Bronze Age and spread across much of Europe, influencing other cultures and lasting until the early Iron Age. [56]

The Urnfield cultures in central Europe are identified primarily from their burial sites, fortified settlements, advancements in metallurgy, and evidence of a stratified society. These sources of material evidence reflect a major cultural and ideological transformation that occurred throughout much of Central and Western Europe. By 1300 BCE, the Urnfield culture had established itself across much of central Europe, including the Rhine River valley. [57]

The latest theories on Urnfield culture migration patterns reflect a shift away from grand narratives of mass migration and instead emphasize complex, regionally diverse dynamics, supported by genetic, isotopic, and archaeological evidence. Isotopic and genomic studies indicate that most Urnfield communities, from Central Europe to the periphery, drew the bulk of their populations locally, rather than through large-scale migrations or abrupt population replacements. [58]

Although mass migration is downplayed, some archaeological sites suggest that individuals of high status (often interpreted as elites or outsiders) could have been highly mobile, reflected both in burials and isotopic, genetic analysis. This suggests the networked movement of select individuals or families rather than wholesale group movements. [59]

The rapid spread of urnfield-type cremation and burial rituals across Europe is now linked to processes of cultural adoption, elite emulation, and regional exchange networks, enabling widespread ideological and cultural shifts without major population turnovers. [60] Studies from Italy, Hungary, and the Carpathian Basin highlight extremely low migration rates within many local urnfield cemeteries, confirming that communities mostly maintained regional genetic continuity with only gradual infiltration from neighbors. [61]

Characteristics of the LRU are documented in the Meuse-Rhine watershed area, especially during the Late Bronze Age. The Urnfield culture was widely distributed across Central and Western Europe, including parts of present-day Netherlands, western Germany, and Belgium, regions that overlap with the Meuse-Rhine watershed. The Lower Rhine urnfield culture developed regional expressions in pottery, burial customs, and settlement patterns, distinguishing it within the broader Urnfield cultural complex.

Intensive mapping projects and archaeological surveys confirm numerous urnfield cemeteries and related settlements in the Dutch, German, and Belgian parts of the eastern Netherlands and the Lower Rhine area. (See sidebar discussion on archaeological mapping). Local densities of urnfields have been recorded along river valleys such as the IJssel, Berkel, and Oude IJssel (tributaries connected to the Meuse-Rhine drainage). The high density of urnfields found along the river valleys highlights the importance of these waterways for the prehistoric communities that lived there. [62]

The concentration of sites suggests that river valleys were a significant feature in the landscape organization and settlement patterns of the urnfield culture. Landscape and settlement analyses indicate that urnfield burial grounds often cluster along sandy ridges, river valleys, and elevated plateaus within the Meuse-Rhine region. [63] 

Settlements in the Meuse–Rhine watershed were typically small-scale, with a relatively dispersed pattern and close connections between burial and habitation sites. Elite hillforts, large fortified settlements, and urban-like centers were more common in southern and eastern Central Europe. Based on archaeological research, the Meuse–Rhine urnfields reflect a transition to smaller, single-family farmsteads, while central European counterparts sometimes maintained larger communal arrangements until later in the period. [64]

The Urnfield culture in the Meuse–Rhine watershed area differs notably from the general Central European Urnfield tradition in burial customs, settlement patterns, cultural boundaries, and socioeconomic dynamics. Drawing on work by Fokkens and later regional studies, these distinctions have been clarified through archaeological, typological, and environmental analyses (re: table two).

Illustration Sixteen: Insights from Archaeological Mapping and Surveys [65]

Click for Larger View | Source: Footnote 66
  • Intensive surveys: This archaeological method involves ‘high-resolution’ fieldwork, with archaeologists systematically walking across a landscape to document all archaeological data, from individual artifacts to architectural remains. This approach is crucial for understanding the finer details of settlement and burial sites.
  • Extensive surveys: In contrast, extensive surveys cover large areas at a ‘lower resolution’ to identify general patterns. These help define the broad distribution of urnfields and other archaeological features, such as the regional concentration along river valleys.
  • Environmental factors: Modern agricultural practices and erosion have significantly impacted urnfield sites. Intensive mapping projects are necessary to identify and protect the remaining evidence, which includes shallow cremation graves and surviving urns.
  • Revealing community structure: By examining the spatial relationship between numerous settlements and the fixed urnfield cemeteries, archaeologists can learn about the social organization and territorial boundaries of Late Bronze and Early Iron Age communities. The discoveries suggest the dissolution of earlier large-scale societal structures into smaller, more autonomous social units. 

Table Two: Comparison of Urnfield Archaeological Characteristics

Archaeological FeatureUnfieldd Culture in
Meuse Rhine
Watershed Area
Unfield Culture in
General Central
Europe
Burial PracticeHybrid, varied forms; democratized cremationMore standardized, elite and communal cremation
Pottery StylesMixed northern/southern, micro-regionalRegional “core” types (e.g. Danuban Lusatian, Bavarian)
Settlement PatternSmall, dispersed, few hillfortsMajor hillforts, cities, concentrated settlements
Cultural BoundariesSharp, micro-regional, pottery-definedPan-regional cultural blocks (pottery/metalwork)
Landscape ImpactIntensive forest clearance, meadow creationRegionally variable, less meadow expansion
Socioeconomic UnitSmaller farms, flexible strategiesLarger communal, persistent settlements
Stratification (buriel evidence)Tribal society divided into autonomous segments led by elders of kin based groups.Rise of warrior elites and class differentiation

The Urnfield phenomena and their local variants in the Meuse-Rhine area are predominantly associated with Y-DNA haplogroup R1b, particularly its branches R1b-L151, R1b-P312, and in some contexts R1b-U106, reflecting the genetic legacy of earlier Bell Beaker and Corded Ware populations in this region.  Ancient DNA from the Meuse-Rhine area (including adjacent Dutch and Belgian urnfield sites) shows a predominance of R1b-L151 lineages, which were already present in the earlier Bell Beaker populations and continued to dominate through the Late Bronze Age and Urnfield period. Subclades such as R1b-P312 (and its further branches DF27, DF19) are well represented, as are R1b-U106, especially toward the Lower Rhine and coastal regions. [66] 

Other haplogroups (I2, G2a, R1a) are present at low levels, but none are as characteristic of the Urnfield burial grounds as R1b-L151-derived types. Most earlier Neolithic haplogroups (like G2a) had largely disappeared from the region’s male gene pool by the Urnfield phase. The Urnfield culture and its local Lower Rhine variant are viewed as carrying forward the patrilineal legacy established by Corded Ware and Bell Beaker expansions, with only minor resurgence of more ancient lineages. The archaeological transition to Urnfield cultural modes in the Meuse-Rhine area shows strong genetic continuity with preceding R1b-dominated groups rather than input from radically new paternal lineages. [67]

Based on genetic studies of ancient DNA, several Y-DNA haplogroups have been associated with the Urnfield culture (1300–750 BCE), primarily identified through archaeological remains in Central Europe. The I2a2b haplogrup dominance suggests continuity from earlier Central European populations, while R1a and R1b subclades reflect Indo-European influences. [68]

From the perspective of ancient DNA, a major challenge in characterizing the Iron Age in Europe has always been cremation’s widespread prevalence. Unlike burials, cremations first destroy most DNA wholesale and then scatter what precious few intact strands might survive.[68a]

Halstatt Culture (~ 800 – 450 BCE)

After the Lower-Rhine Urnfield culture, the Meuse Rhine Watershed area was influenced by the Hallstatt culture during the Early Iron Age (c. 800–450 BCE). The Hallstatt culture is notable for its burial customs, advancements in iron technology and links to early ‘Proto-Celtic’ cultures.

The culture was named after a village in Austria where a significant amount of archaeological remains were discovered. It was a collective set of regional cultures of farmers and metalworkers known for advanced iron and salt production, long-range trade, and elaborate elite burials with unique grave goods that where characterized as having a hierarchical social structure. The Hallstatt culture expanded across wide territories, laid the foundation for later Celtic cultures, and was eventually succeeded by the La Tène culture. [69]

Illustration Seventeen: Map of the Hallstatt Culture

Click for Larger View | Source: Simeon Netchev, Map of the The Hallstatt Culture, published on 22 August 2021, World History Encyclopedia, https://www.worldhistory.org/image/14457/map-of-the-the-hallstatt-culture/

The culture was based on farming, but metal-working was considerably advanced, and by the end of the period long-range trade within the area and with Mediterranean cultures was economically significant. Social distinctions became increasingly important, with emerging elite classes of chieftains and warriors, and perhaps those with other skills. Society is thought to have been organized on a tribal basis, though very little is known about this. Settlement size was generally small, although a few of the largest settlements, like Heuneburg in the south of Germany, evolved into towns rather than villages by modern standards. However, at the end of the period these seem to have been overthrown or abandoned.[70]

The Hallstatt culture is archaeologically divided into four distinct periods known as Hallstatt A, B, C, and D. The four archaeological periods of the Hallstatt culture represent a transition from Late Bronze Age cremation urnfield traditions toward an Early Iron Age Celtic society with evolving burial customs and metalworking technologies.

Hallstatt A and B (c. 1200–800 BCE) are considered part of the Late Bronze Age Urnfield culture. This phase is characterized mainly by cremation burials, with ashes placed in urns, and simple graves. Hallstatt B saw the rise of tumulus (barrow or kurgan) burials becoming common, cremation predominating, and pottery of high craftsmanship. [71]

Hallstatt C and D (c. 800–500 BCE) mark the transition into the Early Iron Age and are traditionally considered the “Hallstatt period” proper and linked with early Celtic culture. Hallstatt C is notable for the first widespread use of iron, with iron swords appearing alongside bronze ones. Both cremation and inhumation occur in this phase. Hallstatt D features mostly inhumation burials, often with daggers instead of swords in western zone graves and changes in pottery and brooch styles. Hallstatt D can be further divided into sub-phases D1–D3 in the western areas, linked mainly to brooch typology differences. Hallstatt D ends around 500 BCE and is succeeded by the La Tène culture. [72]

Table Three: The Four Archaelogical Phases of the General Halstatt Culture [73]

Hallstatt
Period
Date BCE
(approx.)
AgeBurial Type [74]Metal Use
A1200–1000Late Bronze AgeCremation, urn burialBronze; Iron is rare,; characteristic artifacts include bronze tools & ornaments, but none uniquely “Celtic” yet
Hallstatt B1000–800Late Bronze AgeCremation, tumulus burialBronze, some iron;
Continued use of bronze weapons, tools, and vessels.
Brooches and jewelry types begin to diversify, although with regional variation
Hallstatt C800–600 Early Iron AgeCremation and inhumationIron and bronze swords
Notable objects include Hallstatt axes (especially the winged axe), bronze girdle mounts, and long swords..
Early brooch types (“binocular,” “harp,” and “arched”) aid in dating, and distinctive polychrome and unpainted pottery is seen.
Hallstatt D600–500Early Iron AgeMostly inhumationDaggers with horseshoe or “antennae” hilts replace swords in graves, along with elaborately worked brooches that change rapidly in style (snake, drum, and double-drum types).
Other finds include jewelry: armlets, torcs, & ring ornaments, often in bronze or gold for elites.
Elite burials feature imported luxury goods (Greek ceramic vessels, bronze situlae) and grave assemblages indicating extensive trade and high social stratification.
Examples of geometric metal art and occasional animal depictions on objects, especially waterfowl.

Compared to the Alpine core, the Meuse-Rhine Hallstatt region was characterized by more dispersed, rural settlement patterns, lacking centralized fortified hilltop “princely” sites, but still participating in broader cultural traditions through localized elite burial practices and regional economic networks.

The Hallstatt culture was distinguished by a unique combination of social and cultural characteristics which shaped its local and regional identity during the Early Iron Age. Based on archaeological evidence, the core Hallstatt societies exhibited strong social differentiation, with clear evidence of an emerging aristocracy or elite class. Elite burials—often large tumuli with lavish grave goods—signaled the presence of ‘powerful chieftains’ who likely controlled important hilltop settlements and local trade. These hierarchical patterns are reflected in both settlement size and the distribution of grave goods, which indicate a warrior-based leadership and connections between local elites and long-distance networks. [75]

The culture operated within a thriving trade network, exchanging local resources, especially salt, amber, textiles, and agricultural products, for prestige goods such as Mediterranean pottery, wine, and jewelry. This active exchange system transformed both social hierarchies and craft practices and likely linked Meuse-Rhine communities to wider economic and cultural trends of Hallstatt Europe. [76]

Art and traditions shared within the Hallstatt cultural group are considered a foundational, early stage of Celtic culture. The characteristic geometric and linear art style of the Hallstatt period laid the groundwork for later Celtic art. Religious practices, reconstructed from archaeological finds, suggestis nature-oriented beliefs, ritual deposits, and possible ancestor veneration, with customs sometimes differing from central Hallstatt norms. [77]

While sharing core Hallstatt features, societies in the Meuse-Rhine watershed maintained distinct burial customs and patterns of elite display, with strong regional organization and interaction. Studies highlight both integration with wider trade and unique social structures formed through local choices in burial, material culture, and settlement. [78]

Settlement patterns in the Meuse-Rhine Hallstatt region differed markedly from those in the central Alpine Hallstatt core. In the central Alpine heartland, Hallstatt societies featured dense population clusters anchored by fortified hilltop settlements known as “princely seats” or Fürstensitze, such as Heuneburg and Mont Lassois. These sites displayed monumental architecture, richly furnished elite residences, craft workshops, and extensive fortifications, sometimes forming proto-urban agglomerations that could be considered early forms of urbanism north of the Alps. [79]

In contrast, the Meuse-Rhine watershed lacked monumental “princely seats.” Settlements tended to be smaller, less densely clustered, and more dispersed across the landscape. Most communities consisted of rural farmsteads, open villages, or modest hillforts without the same scale of fortification or centralized elite presence found in the Alps. Elite status in the Meuse-Rhine area was showcased more through exceptionally furnished elite burials—often unrelated to any single dominant settlement—rather than through massive settlements or enduring centralized sites. While core Hallstatt sites had significant economic specialization (for example, salt mining at Hallstatt itself and trade-related craft activity at Heuneburg), the Meuse-Rhine zone’s settlements showed less evidence of economic centralization, operating as hubs in more locally oriented agricultural and exchange networks. [80]

The Hallstatt culture eventually declined around 500 BCE, possibly due to the depletion of local salt resources, the emergence of rival trading centers, and shifts in long-range trade routes away from the traditional Hallstatt area. The succeeding La Tène culture became dominant in regional importance. 

Sources:

Feature Banner: The banner at the top of the story features a map of the Meuse and Rhine River watersheds. Overlayed on the map is estimated migratory path of the Griff(is)(es)(ith) paternal YDNA line that is associated with this phylogenetic gap. In addition, various cultures that might have been associated with the generations of ancesors associated with this gap are provided as well as their approximate time periods .

[1] See for example: Peng, Fei (2020). A 13,000-year record of climate- and human-impact-induced flooding in the Lower Meuse. PhD-Thesis – Research and graduation internal, Vrije Universiteit Amsterdam.

Iñigo Olalde, et al., Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups,25 Mar 2025, bioRxiv, https://www.biorxiv.org/content/biorxiv/early/2025/03/25/2025.03.24.644985.full.pdf

[2] See for example: Olson, K. , Krug, E. and Chernyanskii, S. (2025) Natural and Anthropic Environmental Risks to the Rhine River and Delta. Open Journal of Soil Science15, 235-267. doi: 10.4236/ojss.2025.154012.

[3] Scientific Details for haplogroup G-FGC7516:

Click for Larger View | Source: FamilyTreeDNA,Scientific Details on Haplogroup G-FGC7516, https://discover.familytreedna.com/y-dna/G-FGC7516/scientific

The Phylogenetic Gap between G-FGC7516 and G-Z6748 based on images geenrated by the FamilyTreeDNA Globetrekker program.

Click for Larger View | Source: Migratory path of ancestors of G-Y132505, 10 Feb 2025, utilizing FamilyTreeDNA Globe Trekker

[4] See the first part of this story Looking at the Griff(is)(es)(ith) Y-DNA Phylogenetic Gap Associated with the Meuse and Rhine River Watershed – Part OneJ, une 30, 2025.

[5] Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001,  uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

Prehistory of the Netherlands, Wikipedia This page was last edited on 9 July 2025, https://en.wikipedia.org/wiki/Prehistory_of_the_Netherlands

[6] Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review,  https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

Harry Fokkens, Background to Dutch Beakers A Critical Review in  Fokkens, Harry & Franco Nicolis, (eds), Background to Beakers Inquiries into Regional Cultural Backgrounds of the Bell Beaker Complex, Leiden: Sidestone Press, 2012, 9-36

[7] Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

[8] Harry Fokkens, Background to Dutch Beakers A Critical Review in  Fokkens, Harry & Franco Nicolis, (eds), Background to Beakers Inquiries into Regional Cultural Backgrounds of the Bell Beaker Complex, Leiden: Sidestone Press, 2012, 9-36, https://www.sidestone.com/openaccess/9789088900846.pdf

For a similar view, see:

Kliejne, Jos, Embracing Bell Beaker Adopting New Ideas and Objects across Europe during the later 3rd millenium BC (c.2600-200 BC), Lieden: Sidestone Press, 2019

Similar to Fokken, Jos Kleijne’s Embracing Bell Beaker: Adopting New Ideas and Objects across Europe during the later 3rd millennium BC (c. 2600–2000 BC) uses concepts from the archaeology of innovation—like the S-curve, network analysis, and availability models—to explore how social landscapes and mechanisms (including the mobility of objects, people, and ideas) enabled the spread and adaptation of the Bell Beaker phenomenon.

Kliejne includes a comprehensive re-evaluation of how the Bell Beaker phenomenon was adopted and transformed across various European regions. Kleijne analyzes a wide variety of data—including settlement archaeology—to produce a more nuanced view of cultural change, innovation, and local agency during this period. Kleijne

[9] Harry Fokkens, Background to Dutch Beakers A Critical Review, Figure 8, Page 25, https://www.sidestone.com/openaccess/9789088900846.pdf

[10] Harry Fokkens, Background to Dutch Beakers A Critical Review, Pages 24 – 26, https://www.sidestone.com/openaccess/9789088900846.pdf

[11] Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review,  https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

[12] Migrationism and diffusionism, Wikipedia, This page was last edited on 3 September 2025, https://en.wikipedia.org/wiki/Migrationism_and_diffusionism

[13] Armit I, Reich D. The return of the Beaker Folk? Rethinking migration and population change in British prehistory. Antiquity. 2021 Dec;95(384):1464-1477. doi: 10.15184/aqy.2021.129. Epub 2021 Aug 31. PMID: 39524147; PMCID: PMC11550864 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11550864/

Ancient DNA Reveals Impact of the “Beaker Phenomenon” on Prehistoric Europeans, 21 February 2018, Max Planck Institute of Geoanthropology, https://www.shh.mpg.de/842458/beaker-phenomenon

Copper Age Iberians ‘exported’ their culture- but not their genes- all over Europe, 23 Feb 2018, Institute de Biologia Evolutivia, https://www.ibe.upf-csic.es/news/-/asset_publisher/PXTgqZXxlocA/content/id/161929902/maximized#161929902

Olalde, I., Brace, S., Allentoft, M. et al. The Beaker phenomenon and the genomic transformation of northwest Europe. Nature 555, 190–196 (2018). https://doi.org/10.1038/nature25738

Hansen, Victor Davis, Prehistoric pop culture: Deciphering the DNA of the Bell Beaker Complex, 5 Apr 2018, Current Anthropology, https://archaeology.co.uk/articles/features/prehistoric-pop-culture-deciphering-the-dna-of-the-bell-beaker-complex.htm

[14] Olalde I, Altena E, Bourgeois Q, Fokkens H, Amkreutz L, Deguilloux MF, Fichera A, Flas D, Gandini F, Kegler JF, Kootker LM, Leijnse K, Kooijmans LL, Lauwerier R, Miller R, Molthof H, Noiret P, Raemaekers DCM, Rivollat M, Smits L, Stewart JR, Anscher TT, Toussaint M, Callan K, Cheronet O, Frost T, Iliev L, Mah M, Micco A, Oppenheimer J, Patterson I, Qiu L, Soos G, Workman JN, Edwards CJ, Lazaridis I, Mallick S, Patterson N, Rohland N, Richards MB, Pinhasi R, Haak W, Pala M, Reich D. Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups. bioRxiv [Preprint]. 2025 Mar 25:2025.03.24.644985. doi: 10.1101/2025.03.24.644985. PMID: 40196638; PMCID: PMC11974744. (PubMed) https://pubmed.ncbi.nlm.nih.gov/40196638/

[15] S. Arnoldussen, D.J. Huisman, B. van Os, B. Steffens, L. Theunissen, L. Amkreutz, A not so isolated fringe: Dutch later prehistoric (c. 2200 BCE-AD 0) bronze alloy networks from compositional analyses on metals and corrosion layers, Journal of Archaeological Science: Reports, Volume 46, 2022, 103684, ISSN 2352-409X,
https://doi.org/10.1016/j.jasrep.2022.103684 .
(https://www.sciencedirect.com/science/article/pii/S2352409X22003479 )

[16] Bell Beaker Culture, Wikipedia, This page was last edited on 15 September 2025, https://en.wikipedia.org/wiki/Bell_Beaker_culture

See also : Cunliffe, Barry W., The Oxford illustrated prehistory of Europe, Oxford ; New York : Oxford University Press, 1994, 250 – 254, https://archive.org/details/isbn_9780198143857/page/249/mode/2up

[17] Van de Noort, Robert, Exploring Agency Behind the Beaker Phenomenon The navigator’s tale, in Harry Fokkens & Franco Nicolis, eds, Background to Beakers Inquiries in Regional Cultural Backgrounds of the Bell Beaker Complex, Leiden: Slidestone Press, 2012, 61-80, https://www.sidestone.com/books/background-to-beakers

Lanting, J.N. 1997/98, Dates of origin and diffusion of the European logboat, Palaehistoria 39/40, 627-50, https://ugp.rug.nl/Palaeohistoria/article/view/25107/22563 ; also https://www.ancientportsantiques.com/wp-content/uploads/Documents/ETUDESarchivees/Navires/Documents/Lanting2000-Logboats.pdf

See also : Lanting J.N. and Van der Waals, J. D. 1976 Beaker culture relations in the Lower Rhine Basin, in Lanting J.N. and Van der Waals, J.D. (eds. Glockenbecher Symosium Oberried 1974, Bussum: Fibula – Van Dishoech, 1-80

[18] Fokens, H. , B.J.W. Steffens & S.F.M. van As, Farmers, fishers, fowlers, hunters Knowledge generated by development-led archaeology about the Late Neothithic, the Early Bronze Age and the start of the Middle Bronze Age (2850 – 1500 cal BC) in the Netherlands, Amersfoort: Rijksdienst voor het Cultureel Erfgoed, 2016, Page 288 , https://cultureelerfgoed.info/download/pdf/NAR053_Farmers_fishers_fowlers_hunters.pdf

[19] Fokens, H. , B.J.W. Steffens & S.F.M. van As, Farmers, fishers, fowlers, hunters Knowledge generated by development-led archaeology about the Late Neothithic, the Early Bronze Age and the start of the Middle Bronze Age (2850 – 1500 cal BC) in the Netherlands, Amersfoort: Rijksdienst voor het Cultureel Erfgoed, 2016, Page 292, https://cultureelerfgoed.info/download/pdf/NAR053_Farmers_fishers_fowlers_hunters.pdf

[20] Dugout canoe, Wikipedia, This page was last edited on 15 June 2025, https://en.wikipedia.org/wiki/Dugout_canoe

Pesse canoe, Wikipedia, This page was last edited on 5 September 2025, https://en.wikipedia.org/wiki/Pesse_canoe

[21] Henk J.A Berendsen, Esther Stouthamer, Late Weichselian and Holocene palaeogeography of the Rhine–Meuse delta, The Netherlands, Palaeogeography, Palaeoclimatology, Palaeoecology, Volume 161, Issues 3–4, 2000, Pages 311-335, ISSN 0031-0182, https://doi.org/10.1016/S0031-0182(00)00073-0 .
(https://www.sciencedirect.com/science/article/pii/S0031018200000730 )

Lanting, J.N. 1997/98, Dates of origin and diffusion of the European logboat, Palaehistoria 39/40, 627-50, https://ugp.rug.nl/Palaeohistoria/article/view/25107/22563 ; also https://www.ancientportsantiques.com/wp-content/uploads/Documents/ETUDESarchivees/Navires/Documents/Lanting2000-Logboats.pdf

Gouw, M. J. P. and Hijma, M. P.: From apex to shoreline: fluvio-deltaic architecture for the Holocene Rhine–Meuse delta, the Netherlands, Earth Surf. Dynam., 10, 43–64, https://doi.org/10.5194/esurf-10-43-2022 , 2022

[22] Pesse canoe, Wikipedia, This page was last edited on 5 September 2025, https://en.wikipedia.org/wiki/Pesse_canoe

[23] Lanting, J.N. 1997/98, Dates of origin and diffusion of the European logboat, Palaehistoria 39/40, 627-50, https://ugp.rug.nl/Palaeohistoria/article/view/25107/22563 ; also https://www.ancientportsantiques.com/wp-content/uploads/Documents/ETUDESarchivees/Navires/Documents/Lanting2000-Logboats.pdf

[24] Lanting, J.N. 1997/98, Dates of origin and diffusion of the European logboat, Palaehistoria 39/40, 627-50, https://ugp.rug.nl/Palaeohistoria/article/view/25107/22563 ; also https://www.ancientportsantiques.com/wp-content/uploads/Documents/ETUDESarchivees/Navires/Documents/Lanting2000-Logboats.pdf

[25] Van de Noort, Robert, Exploring Agency Behind the Beaker Phenomenon The navigator’s tale, in Harry Fokkens & Franco Nicolis, eds, Background to Beakers Inquiries in Regional Cultural Backgrounds of the Bell Beaker Complex, Leiden: Slidestone Press, 2012, 61-80, https://www.sidestone.com/books/background-to-beakers

[26] Harry Fokkens, Background to Dutch Beakers A Critical Review in  Fokkens, Harry & Franco Nicolis, (eds), Background to Beakers Inquiries into Regional Cultural Backgrounds of the Bell Beaker Complex, Leiden: Sidestone Press, 2012, 9-36

Kliejne, Jos, Embracing Bell Beaker Adopting New Ideas and Objects across Europe during the later 3rd millenium BC (c.2600-200 BC), Lieden: Sidestone Press, 2019

Dunkley, Mark Travelling by water: A chronology of prehistoric boat archaeology/mobility in England, no date, Ancient Coastal Settlements, Ports and Harbours, https://www.ancientportsantiques.com/wp-content/uploads/Documents/ETUDESarchivees/Navires/Documents/Dunkley2013-PrehistoricBoats.pdf

Van de Noort, Robert, Exploring Agency Behind the Beaker Phenomenon The navigator’s tale, in Harry Fokkens & Franco Nicolis, eds, Background to Beakers Inquiries in Regional Cultural Backgrounds of the Bell Beaker Complex, Leiden: Slidestone Press, 2012, 61-80, https://www.sidestone.com/books/background-to-beakers

Quentin P. J. Bourgeois et al. Spatiotemporal reconstruction of Corded Ware and Bell Beaker burial rituals reveals complex dynamics divergent from steppe ancestry. Sci. Adv. 11, eadx2262 (2025) DOI:10.1126/sciadv.adx2262

[27] Modification of a map originally in Harry Fokkens, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001, uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

[28] Elp Culture, Wikipedia, This page was last edited on 17 October 2024, https://en.wikipedia.org/wiki/Elp_culture

Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001, uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

[29] Elp Culture, Wikipedia, This page was last edited on 17 October 2024, https://en.wikipedia.org/wiki/Elp_culture

Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001, uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

[30] Elp Culture, Wikipedia, This page was last edited on 17 October 2024, https://en.wikipedia.org/wiki/Elp_culture

[31] The Tumulus Culture ” was the descendant of the Unetice culture. Its heartland was the area previously occupied by the Unetice culture, and its territory included parts of Germany, the Czech Republic, Austria, Switzerland, the Carpathian Basin, Poland and France. It was succeeded by the Late Bronze Age Urnfield culture and part of the origin of the Italic and Celtic cultures.”

Tumulus Culture, Wikipedia, This page was last edited on 2 July 2025, https://en.wikipedia.org/wiki/Tumulus_culture

[32] Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001, uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

Elp Culture, Wikipedia, This page was last edited on 17 October 2024, https://en.wikipedia.org/wiki/Elp_culture

Tumulus Culture, Wikipedia, This page was last edited on 2 July 2025, https://en.wikipedia.org/wiki/Tumulus_culture

[33] Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001, uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

Fokens, H. , B.J.W. Steffens & S.F.M. van As, Farmers, fishers, fowlers, hunters Knowledge generated by development-led archaeology about the Late Neothithic, the Early Bronze Age and the start of the Middle Bronze Age (2850 – 1500 cal BC) in the Netherlands, Amersfoort: Rijksdienst voor het Cultureel Erfgoed, 2016, Page 38 https://collectie.huisvanhilde.nl/pdf/rce_nar-053_farmers_fishers_fowlers_hunters.pdf

Nicolay, J. (2010). “Settlement research and material culture in the northern Netherlands: Herrenhöfe and other evidence of socio-political differentiation.” Settlement and Coastal Research in the Southern North Sea Region, 33, 119-132.

Meier, D. (2013). “Settlements and sediments: The cultural landscape of the North Sea coastal zone.” The Oxford Handbook of Wetland Archaeology, Oxford University Press, 473-489.

Zimmermann, W. H., “Why was cattle-stalling introduced in prehistory? The significance of byre and stable.” Charlottee Fabech & Jytte Ringtved, eds, Settlement and Landscape, Aarhus University Press, 1999, 301-318. https://www.researchgate.net/publication/264478571_Why_was_cattle-stalling_introduced_in_prehistory_The_significance_of_byre_and_stable_and_of_outwintering

Armstrong Oma, K. (2013). “Human-animal relationships: Mutual becomings in the household of Scandinavia and Sicily 900-500 BC.” Anthropology & Archaeology, 2013, Oslo Academic Press.

[34] Hilversum Culture, Wikipedia This page was last edited on 8 July 2025, https://en.wikipedia.org/wiki/Hilversum_culture

Patterson, N. et al. Large-scale migration into Britain during the Middle to Late Bronze Age. Nature, 601(7894), 2002, pp. 588-594. (doi: 10.1038/s41586-021-04287-4), https://eprints.gla.ac.uk/263057/2/263057.pdf

Högberg, Anders. (2011). Peter Clark, ed., Bronze Age Connections: Cultural Contact in Prehistoric Europe. European Journal of Archaeology. 14. 304-306. 10.1179/eja.2011.14.1-2.304

Morris, Francis Michael, Cross-North Sea Contacts in the Roman Period, OXFORD Journal of Archaeology 34(4) 415–438 2015, 415- 438, https://www.researchgate.net/publication/282844822_Cross-North_Sea_Contacts_in_the_Roman_Period

[35] Fokkens, Harry, The longhouse as a central element in Bronze Age daily life. In J. Bourgeois, I. Bourgeois, & B. Charetté (Eds.), Bronze Age and Iron Age communities in North-Western Europe , 2005, pp. 9-38. Brussels. Retrieved from https://hdl.handle.net/1887/19826

Arnoldusse, Stijn, A Living Landscape Bronze Age Settlement Sites in the Dutch River area (c 2000-800 BC), Lieden: Sidestone Press, 2008, https://www.sidestone.com/openaccess/9789088900105.pdf

[36] Fokkens, H. The longhouse as a central element in Bronze Age daily life. In J. Bourgeois, I. Bourgeois, & B. Charetté (Eds.), Bronze Age and Iron Age communities in North-Western Europe (pp. 9-38). Brussels. Retrieved from https://hdl.handle.net/1887/19826  

[37] Fokkens, H. The longhouse as a central element in Bronze Age daily life. In J. Bourgeois, I. Bourgeois, & B. Charetté (Eds.), Bronze Age and Iron Age communities in North-Western Europe (pp. 9-38). Brussels. Retrieved from https://hdl.handle.net/1887/19826  

Arnoldusse, Stijn, A Living Landscape Bronze Age Settlement Sites in the Dutch River area (c 2000-800 BC), Lieden: Sidestone Press, 2008, https://www.sidestone.com/openaccess/9789088900105.pdf

[38] Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001,  uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

[39] Tumulus Culture, Wikipedia, This page was last edited on 2 July 2025, https://en.wikipedia.org/wiki/Tumulus_culture

[40] Arnoldussen Stijn and Harry Fokkens, Bronze Age settlements in the Low Countries: an overview, Chapter Two, in Arnoldussen, Stijn and Harry Fokkens, eds, Bronze Age Settlements in the Low Countries, Oxford, UK: Oxbow Books, 2008, 17-40, https://www.researchgate.net/publication/349414629_Bronze_Age_settlements_in_the_Low_Countries_an_overview#:~:text=German%20Urnfield%20Culture.-,In%20the,showing%20influence%20of%20the%20Southern

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Rowlett, Elsebeth and Ralph, Champagne and the Ardennes During the Bronze and Iron Ages, MSSE Annual of the Museum of Art and Archeology, Number 5, 1971 University of Missouri-Columbia, 23 – 25 , https://maa.missouri.edu/sites/default/files/file-uploads/2021-12/muse_1971_-_vol._5_optimized.pdf

Quiles, Carlos, The origins of the Tumulus culture: Proto-Lusatian and potential Proto-Balto-Slavic origins, 29 Mar 2019, Indo-European.eu, https://indo-european.eu/2018/03/the-origins-of-the-tumulus-culture-proto-lusatian-and-potential-proto-balto-slavic-origins/

[41] Tumulus Culture, Wikipedia, This page was last edited on 2 July 2025, https://en.wikipedia.org/wiki/Tumulus_culture

Quiles, Carlos, The origins of the Tumulus culture: Proto-Lusatian and potential Proto-Balto-Slavic origins, 29 Mar 2019, Indo-European.eu, https://indo-european.eu/2018/03/the-origins-of-the-tumulus-culture-proto-lusatian-and-potential-proto-balto-slavic-origins/

Harding A. Bronze Age Encounters: Violent or Peaceful? In: Horn C, Kristiansen K, eds. Warfare in Bronze Age Society. Cambridge University Press; 2018: 16-22.

LP Louw e Kooijmans, An Early/Middle Bronze Age multiple burial at Wassenaar, the Netherlands, Analecta Praehisotorica Leidensia, 26, 1 – 20, https://scholarlypublications.universiteitleiden.nl/access/item%3A2732058/view

Fontijn, D. and Fokkens, H., ‘The emergence of Early Iron Age “chieftains’ graves” in the southern Netherlands. Reconsidering transformations in burial and depositional practices’, in Haselgrove, C. and Pope, R. (eds.), The Earlier Iron Age in Britain and the Near Continent (Oxford: Oxbow Books, 2007), pp. 354–373 , https://www.researchgate.net/publication/333124942_The_emergence_of_Early_Iron_Age_%27chieftains%27_graves%27_in_the_southern_Netherlands_reconsidering_transformations_in_burial_and_depositional_practices

[42] Fontijn, D. and Fokkens, H., ‘The emergence of Early Iron Age “chieftains’ graves” in the southern Netherlands. Reconsidering transformations in burial and depositional practices’, in Haselgrove, C. and Pope, R. (eds.), The Earlier Iron Age in Britain and the Near Continent (Oxford: Oxbow Books, 2007), pp. 354–373 , https://www.researchgate.net/publication/333124942_The_emergence_of_Early_Iron_Age_%27chieftains%27_graves%27_in_the_southern_Netherlands_reconsidering_transformations_in_burial_and_depositional_practices

Pope, Rachael, and Colin Haselgrove, The Earlier Iron Age in Britain and the Near Continent, Oxford: Oxbrow, 2006

Warming, Rolf Fabricius ,ed, Violence and Warfare in Social Context Archaeological and Historical Studies, Stockholm: Stockholm University, 2025 https://www.diva-portal.org/smash/get/diva2:1947235/FULLTEXT01.pdf

For warfare in the Neolithic, see:

Christensen , Jonas, Warfare in the Neolithic , Acta Archaeologica vol. 75, 2004, pp. 129–156 , https://faculty.uml.edu/ethan_spanier/Teaching/documents/WarfareintheNeolithic.pdf

[43] Gerritsen, Fokke, Local Identities : Landscape and Community in the Late Prehistoric Meuse-Demer-Scheldt Region, Amersterdam Archaeological Studies, 9, Amsterdam: Amsterdam University Press, 2003, https://www.academia.edu/451425/Local_Identities_Landscape_and_Community_in_the_Late_Prehistoric_Meuse_Demer_Scheldt_Region

Quiles, Carlos, The origins of the Tumulus culture: Proto-Lusatian and potential Proto-Balto-Slavic origins, 29 Mar 2019, Indo-European.eu, https://indo-european.eu/2018/03/the-origins-of-the-tumulus-culture-proto-lusatian-and-potential-proto-balto-slavic-origins/

[44] Tumulus Culture, Wikipedia, This page was last edited on 2 July 2025, https://en.wikipedia.org/wiki/Tumulus_culture

Quiles, Carlos, The origins of the Tumulus culture: Proto-Lusatian and potential Proto-Balto-Slavic origins, 29 Mar 2019, Indo-European.eu, https://indo-european.eu/2018/03/the-origins-of-the-tumulus-culture-proto-lusatian-and-potential-proto-balto-slavic-origins/

Gerritsen, Fokke, Local Identities : Landscape and Community in the Late Prehistoric Meuse-Demer-Scheldt Region, Amersterdam Archaeological Studies, 9, Amsterdam: Amsterdam University Press, 2003, https://www.academia.edu/451425/Local_Identities_Landscape_and_Community_in_the_Late_Prehistoric_Meuse_Demer_Scheldt_Region

Falkenstein, Frank, The Development of Burial Rites from the Tumulus to the Urnfield Culture in Southern Central Europe, 2011, E. Borgna/S. Müller-Celka (eds.), Ancestral Landscapes. Burial Mounds in the Copper and Bronze Ages (Central and Eastern Europe – Balkans – Adriatic – Aegean, 4th-2nd millenium B.C.). Proceedings of the International Conference held in Udine, May 5th-18th 2008, https://www.academia.edu/5438219/The_Development_of_Burial_Rites_from_the_Tumulus_to_the_Urnfield_Culture_in_Southern_Central_Europe

Schnieder, Seth , Ancestor Veneration and Ceramic Curation: An Analysis from Speckhau Tumulus 17, Southwest Germany, Masters Thesis, University of Wisconsin, 2003, https://www.academia.edu/731220/Ancestor_Veneration_and_Ceramic_Curation_An_Analysis_from_Speckhau_Tumulus_17_Southwest_Germany

Sasjavdv, Early Iron Age (800–500 BC) elite burials in the Low Countries, 17 Dec 2019, Fragmenting the Chieftain, http://vandervaart-verschoof.com/fragmenting-the-chieftain-the-elite-burials-of-the-early-iron-age-800-500-bc-in-the-low-countries/

[45] Capuzzo G, De Mulder G, Sabaux C, et al. Final Neolithic and Bronze Age Funerary Practices and Population Dynamics in Belgium, The Impact of Radiocarbon Dating Cremated Bones, Radiocarbon. 2023;65(1):51-80. doi: https://10.1017/RDC.2022.94  

Holst, Mads Kähler, ‘Burials’, in Harry Fokkens, and Anthony Harding (eds), The Oxford Handbook of the European Bronze Age (2013; online edn, Oxford Academic, 5 Sept. 2013), https://doi.org/10.1093/oxfordhb/9780199572861.013.0006

Faukenstein, Frank, The Development of Burial Rites from the Tumulus to the Urnfield Culture in Sourthern Central Europe, in Elisabetta Borgna and Sylvie Mülle Celka, Ancestral Landscapes. Lyon: Maison de l’orient et de la Méditerranée, 2011, 229 – 340, https://www.academia.edu/5438219/The_Development_of_Burial_Rites_from_the_Tumulus_to_the_Urnfield_Culture_in_Southern_Central_Europe

De Mulder G, van Strydonck M, Boudin M. The Impact of Cremated Bone Dating on the Archaeological Chronology of the Low Countries. Radiocarbon. 2009;51(2):579-600. doi: https://10.1017/S0033822200055946 

De Mulder G, van Strydonck M, De Clercq W. 14C Dating of “Brandgrubengräber” from the Bronze Age to the Roman Period in Western Flanders (Belgium). Radiocarbon. 2013;55(3):1233-1245. doi: https://10.1017/S0033822200048141 

[46] Faukenstein, Frank, The Development of Burial Rites from the Tumulus to the Urnfield Culture in Sourthern Central Europe, in Elisabetta Borgna and Sylvie Mülle Celka, Ancestral Landscapes. Lyon: Maison de l’orient et de la Méditerranée, 2011, 229 – 340, https://www.academia.edu/5438219/The_Development_of_Burial_Rites_from_the_Tumulus_to_the_Urnfield_Culture_in_Southern_Central_Europe

[47] Holst, Mads Kähler, ‘Burials’, in Harry Fokkens, and Anthony Harding (eds), The Oxford Handbook of the European Bronze Age (2013; online edn, Oxford Academic, 5 Sept. 2013), https://doi.org/10.1093/oxfordhb/9780199572861.013.0006

[47] Tumulus culture, Wikipedia, https://en.wikipedia.org/wiki/Tumulus_culture

Quiles, Carlos, The origins of the Tumulus culture: Proto-Lusatian and potential Proto-Balto-Slavic origins, 29 Mar 2018, Indo-European.eu, https://indo-european.eu/2018/03/the-origins-of-the-tumulus-culture-proto-lusatian-and-potential-proto-balto-slavic-origins/

[48] Falkenstein, The Development of Burial Rites from the Tumulus to the Urnfield Culture in Southern Central Europe

Capuzzo G, De Mulder G, Sabaux C, et al. Final Neolithic and Bronze Age Funerary Practices and Population Dynaics in Belgium, The Ipact of Radiocarbon Dating Cremated Bones,  Radiocarbon. 2023;65(1):51-80. doi: https://10.1017/RDC.2022.94

Holst, Mads Kähler, ‘Burials’, in Harry Fokkens, and Anthony Harding (eds), The Oxford Handbook of the European Bronze Age (2013; online edn, Oxford Academic, 5 Sept. 2013), https://doi.org/10.1093/oxfordhb/9780199572861.013.0006

[49] See for example Cavazzuti, C., Horváth, A., Gémes, A. et al. Isotope and archaeobotanical analysis reveal radical changes in mobility, diet and inequalities around 1500 BCE at the core of Europe.Sci Rep 15, 17494 (2025). https://doi.org/10.1038/s41598-025-01113-z

[50] Urnfield culture, Wikipedia, This page was last edited on 4 April 2025,, https://en.wikipedia.org/wiki/Urnfield_culture

Riedinger, Edward A., Urnfield Culture, 2022, EBSCO, https://www.ebsco.com/research-starters/anthropology/urnfield-culture

Urnfield culture, Distributed Wikipedia, May 2016, https://nzt-eth.ipns.dweb.link/wiki/Urnfield_culture.html

Smith, M. A., A Study in Urnfield Interpretations in Middle Europe. Zephyrvs8. 2009, https://revistas.usal.es/uno/index.php/0514-7336/article/view/3640MÁS

[51] Hugh McColl, Guus Kroonen, Thomaz Pinotti, William Barrie, John Koch, Johan Ling, Jean-Paul Demoule, Kristian Kristiansen, Martin Sikora, Eske Willerslev, Tracing the spread of Celtic languages using ancient genomics, bioRxiv 2025.02.28.640770; doi: https://doi.org/10.1101/2025.02.28.640770

Bretos Ezcurra, M., Rohrlach, A.B., Papac, L. et al. Genomic insights from a final Bronze Age community buried in a collective tumulus in an Urnfield settlement in Northeastern Iberia. Commun Biol 8, 1299 (2025). https://doi.org/10.1038/s42003-025-08668-7

[52] Fokkens, Harry, The genesis of urnfields: Economic crisis or ideological change?, June 1997, Antiquity, 71 (272), DOI:10.1017/S0003598X00084970 , https://www.researchgate.net/publication/28646976_The_genesis_of_urnfields_Economic_crisis_or_ideological_change

[53] Arjan Louwen, Breaking and Making the Ancestors: Piecing together the urnfield mortuary process in the Lower-Rhine-Basin, ca. 1300 – 400 BC, Leiden, Slidestone Press, 2021, Page 12, https://www.academia.edu/66691895/Breaking_and_Making_the_Ancestors_Piecing_together_the_urnfield_mortuary_process_in_the_Lower_Rhine_Basin_ca_1300_400_BC?uc-g-sw=23584249

[54] Fokkens, Harry, The genesis of urnfields: Economic crisis or ideological change?, June 1997, Antiquity, 71 (272), DOI:10.1017/S0003598X00084970 , https://www.researchgate.net/publication/28646976_The_genesis_of_urnfields_Economic_crisis_or_ideological_change

[55] Hand drawn figure is from Fig. 3.11: Schematic overview of the types of archaeological features associated with urnfield graves and their terminology Page 61 in A.J. Louwen, Breaking and making the ancestors. piecing together the urnfield
mortuary process in the Lower-Rhine-Basin, ca. 1300–400 BC, 17 Aug 2021, PhD Thesis, Leiden: Sidestone Press, https://hdl.handle.net/1887/3185517

[56] Prior to the Urnfield-type cremation practices of the Late Bronze Age, burial practices in Europe varied significantly by region and time period, though many centered on inhumation under burial mounds or in communal tombs. A shift from these inhumation practices to widespread cremation defines the start of the Urnfield period in many areas. In the Neolithic period, burial customs across Europe were diverse and often involved collective burials in large monuments rather than individual graves. 

Unlike the collective burials of the Neolithic, the Middle Bronze Age is characterized by the widespread use of single burials. The dead were buried in barrows, or tumuli—mounds of earth and stones raised over one or more graves. Some of these mounds were initially built for a primary, often high-status, burial and then reused for subsequent interments.

During the later Tumulus culture, cremation began to appear alongside inhumation. In some instances, cremated remains were interred in barrows as secondary burials, foreshadowing the later Urnfield practices. 

See:

Fowler, Chris, and Chris Scarre, ‘Mortuary Practices and Bodily Representations in North-West Europe’,in Chris Fowler, Jan Harding, and Daniela Hofmann (eds), The Oxford Handbook of Neolithic Europe(2015; online edn, Oxford Academic, 5 Dec. 2014), https://doi.org/10.1093/oxfordhb/9780199545841.013.054 

Tumulus Clture, Wikipedia, This page was last edited on 2 July 2025, https://en.wikipedia.org/wiki/Tumulus_culture

Riedinger, Edward A., Urnfield Culture, EBSCO Knowledge advantage, , 2022, https://www.ebsco.com/research-starters/anthropology/urnfield-culture

“Late Bronze Age Urnfields of Central Europe .” Ancient Europe, 8000 B.C. to A.D. 1000: Encyclopedia of the Barbarian World. . Encyclopedia.com. (September 2, 2025). https://www.encyclopedia.com/humanities/encyclopedias-almanacs-transcripts-and-maps/late-bronze-age-urnfields-central-europe

[57] Urnfield culture, Wikipedia, This page was last edited on 4 April 2025,, https://en.wikipedia.org/wiki/Urnfield_culture

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[58] Giacomo Capuzzo, Elisavet Stamataki, Michael Allen Beck De Lotto, Silvia Pettarin, Philippe Claeys, Nadine Mattielli, Giovanni Tasca, Christophe Snoeck, A multi-proxy approach to reconstruct chronology, human mobility, and funerary practices at the Late Bronze-Early Iron Age urnfield of San Valentino (San Vito al Tagliamento, Italy), November 7, 2024, PLOS https://doi.org/10.1371/journal.pone.0309649 , https://journals.plos.org/plosone/article?id=10.1371%2Fjournal.pone.0309649

Claudio Cavazzuti, Tamás Hajdu, Federico Lugli, Alessandra Sperduti, Magdolna Vicze, Aniko Horváth, István Major, Mihály Molnár, László Palcsu, Viktória Kiss, Human mobility in a Bronze Age Vatya ‘urnfield’ and the life history of a high-status woman, July 28, 2021, PLOS, https://doi.org/10.1371/journal.pone.0254360, https://journals.plos.org/plosone/article?id=10.1371%2Fjournal.pone.0254360

[59] Claudio Cavazzuti, et al, Human mobility in a Bronze Age Vatya ‘urnfield’ and the life history of a high-status woman, July 28, 2021, PLOS, https://doi.org/10.1371/journal.pone.0254360, https://journals.plos.org/plosone/article?id=10.1371%2Fjournal.pone.0254360

Frank AB, May J, Sabatini S, Schopper F, Frei R, Kaul F, Storch S, Hansen S, Kristiansen K, Frei KM. A Late Bronze Age foreign elite? Investigating mobility patterns at Seddin, Germany. PLoS One. 2025 Sep 10;20(9):e0330390. doi: 10.1371/journal.pone.0330390. PMID: 40929027; PMCID: PMC12422465 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC12422465/

[60] Renan Falcheti Peixoto and Francesco Iacono, Urnfield Bronze Connections: Rethinking Late Bronze Age Mobility, Ocnus 30 (2022): 149-172; doi: 10.12876/OCNUS3010; ISSN 1122-6315; 149-165, https://www.academia.edu/97794355/Urnfield_Bronze_Connections_Rethinking_Late_Bronze_Age_Mobility

Giacomo Capuzzo, et al, A multi-proxy approach to reconstruct chronology, human mobility, and funerary practices at the Late Bronze-Early Iron Age urnfield of San Valentino (San Vito al Tagliamento, Italy), November 7, 2024, PLOS https://doi.org/10.1371/journal.pone.0309649 , https://journals.plos.org/plosone/article?id=10.1371%2Fjournal.pone.0309649

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[61] This picture of stability is often reconstructed by combining genetic data with isotopic analysis, which reveals the geographical origins of individuals based on the chemical composition of their remains. These genetic and isotopic findings indicate that the spread of the Urnfield cultural “package,” which included cremation burial in urns, does not necessarily correlate with large-scale population migration. Instead, it suggests a process of cultural transmission where the burial custom was adopted by existing local communities across Europe. 

San Valentino: Research on the Urnfield cemetery at San Valentino in northeastern Italy, published in 2024, used strontium isotope analysis to conclude that the people buried there were a local community who exploited nearby resources.

Northern vs. Central/Southern Italy: Studies on Celtic migrations in the Iron Age found that, while northern Italy experienced biological admixture with new groups, central and southern Italy showed a strong, “pre-Iron Age” genetic background. This indicates that the local substrate YDNA composition remained very stable through the end of the Bronze Age and into the Iron Age in some areas. 

See:

Piccirilli E, Sorrentino R, Lugli F, Bortolini E, Silvestrini S, Cavazzuti C, Conti S, Czifra S, Gyenesei K, Köhler K, Tankó K, Vazzana A, Jerem E, Cipriani A, Gottarelli A, Belcastro MG, Hajdu T, Benazzi S. New insights on Celtic migration in Hungary and Italy through the analysis of non-metric dental traits. PLoS One. 2023 Oct 18;18(10):e0293090. doi: 10.1371/journal.pone.0293090. Erratum in: PLoS One. 2024 Dec 26;19(12):e0316684. doi: 10.1371/journal.pone.0316684. PMID: 37851635; PMCID: PMC10584115, https://pmc.ncbi.nlm.nih.gov/articles/PMC10584115/

Capuzzo G, Stamataki E, Beck De Lotto MA, Pettarin S, Claeys P, Mattielli N, Tasca G, Snoeck C. A multi-proxy approach to reconstruct chronology, human mobility, and funerary practices at the Late Bronze-Early Iron Age urnfield of San Valentino (San Vito al Tagliamento, Italy). PLoS One. 2024 Nov 7;19(11):e0309649. doi: 10.1371/journal.pone.0309649. PMID: 39509443; PMCID: PMC11542862 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11542862/; also https://journals.plos.org/plosone/article?id=10.1371%2Fjournal.pone.0309649

Cavazzuti, C., Horváth, A., Gémes, A. et al. Isotope and archaeobotanical analysis reveal radical changes in mobility, diet and inequalities around 1500 BCE at the core of Europe.Sci Rep 15, 17494 (2025). https://doi.org/10.1038/s41598-025-01113-z

Carpathian Basin: Research on populations from the Bronze Age through the Early Copper Age in the Carpathian Basin revealed a high degree of genetic continuity, despite changes in material culture and burial practices. Some contemporaneous communities showed radically different social structures, ranging from genetically diverse and open to more homogenous and closed.

Western Hungary: A 2023 study focusing on Bronze Age communities in western Hungary used ancient DNA to confirm the genetic continuity of some groups, such as the Kisapostag population, which contributed to the genetic basis of the succeeding Encrusted Pottery culture. The findings also highlighted evidence of patrilocality and social stability.

Archaeogenetic limitations: Scholars note that cremation, the defining Urnfield burial rite, destroys DNA, which makes direct genetic analysis of cremated individuals extremely difficult. As a result, many conclusions about Urnfield population genetics are based on isotopic data from cremated remains and genetic analysis of coeval inhumation burials. 

See:

Capuzzo G, Stamataki E, Beck De Lotto MA, Pettarin S, Claeys P, Mattielli N, Tasca G, Snoeck C. A multi-proxy approach to reconstruct chronology, human mobility, and funerary practices at the Late Bronze-Early Iron Age urnfield of San Valentino (San Vito al Tagliamento, Italy). PLoS One. 2024 Nov 7;19(11):e0309649. doi: 10.1371/journal.pone.0309649. PMID: 39509443; PMCID: PMC11542862 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11542862/; also https://journals.plos.org/plosone/article?id=10.1371%2Fjournal.pone.0309649

Cavazzuti, C., Arena, A., Cardarelli, A. et al. The First ‘Urnfields’ in the Plains of the Danube and the Po. J World Prehist 35, 45–86 (2022). https://doi.org/10.1007/s10963-022-09164-0

Gerber D, Szeifert B, Székely O, Egyed B, Gyuris B, Giblin JI, Horváth A, Köhler K, Kulcsár G, Kustár Á, Major I, Molnár M, Palcsu L, Szeverényi V, Fábián S, Mende BG, Bondár M, Ari E, Kiss V, Szécsényi-Nagy A. Interdisciplinary Analyses of Bronze Age Communities from Western Hungary Reveal Complex Population Histories. Mol Biol Evol. 2023 Sep 1;40(9):msad182. doi: 10.1093/molbev/msad182. PMID: 37562011; PMCID: PMC10473862., https://pmc.ncbi.nlm.nih.gov/articles/PMC10473862/

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Riedinger, Edward A., Urnfield Culture, EBSCO Knowledge advantage, 2022, urnfield-culture

Theuws, Frans, and Nico Roymans, eds. Land and Ancestors: Cultural Dynamics in the Urnfield Period and the Middle Age in the Southern Netherlands. Amsterdam: University Press, 1999. Several chapters in this work deal with Urnfield culture in the late Bronze and early Iron Ages at its further extensions in northwestern Europe.

Urnfield Culture, Wikipedia, This page was last edited on 2 September 2025, https://en.wikipedia.org/wiki/Urnfield_culture

Roymans, N.G.A.M. ; Kortlang, F.P. / Urnfield symbolism, ancestors and the land in the Lower Rhine Region. Land and ancestors. Cultural dynamics in the Urnfield period and the Middle Ages in the Southern Netherlands. editor / F. Theuws ; N.G.A.M. Roymans. Amsterdam, 1999. pp. 33-62 (Amsterdam archaeological studies; 4). 

Fokkens, H., & Arnoldussen, S. (2008). Towards new models. In F. H. Arnoldussen A. (Ed.), Bronze Age settlements in the Low Countries (pp. 1-16). Oxford: Oxbow books. Retrieved from https://hdl.handle.net/1887/13819

Fokkens H. The genesis of urnfields: economic crisis or ideological change? Antiquity. 1997;71(272):360-373. doi:10.1017/S0003598X00084970, https://www.researchgate.net/publication/28646976_The_genesis_of_urnfields_Economic_crisis_or_ideological_change

Helene Agerskov Rose, Lisbeth Christensen and Arjan Louwen, eds, Beyond Urnfields New Perspectives on Late Bronze Age – Early Iron Age Funerary Practices in Northwest Europe,  Keil: Verlag Ludwig 2003 , https://doi.org/10.2369/9783869354439.19-louwen , https://www.verlag-ludwig.de/files/10_2369_9783869354439_19_louwen.pdf

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Fokkens, H., & Arnoldussen, S. (2008). Towards new models. In F. H. Arnoldussen A. (Ed.), Bronze Age settlements in the Low Countries (pp. 1-16). Oxford: Oxbow books. Retrieved from https://hdl.handle.net/1887/13819

Roy van Beek and Arjan Louwen, The centrality of urnfields. Second thoughts on structure and stability  of Late Bronze Age and Early Iron Age cultural landscapes in the Low  Countries, Fontijn, D., A.J. Louwen, S. van der Vaart & K. Wentink (eds) 2013:  Beyond Barrows. Current research on the structuration and  perception of the Prehistoric Landscape through Monuments.  Leiden: Sidestone Press, 2013, https://www.academia.edu/3160119/Beek_R_van_A_Louwen_2013_The_centrality_of_urnfields_Second_thoughts_on_structure_and_stability_of_Late_Bronze_Age_and_Early_Iron_Age_cultural_landscapes_in_the_Low_Countries_in_Fontijn_D_A_J_Louwen_S_van_der_Vaart_K_Wentink_eds_Beyond_Barrows_Leiden_81_112

Roymans, N.G.A.M. ; Kortlang, F.P. / Urnfield symbolism, ancestors and the land in the Lower Rhine Region. Land and ancestors. Cultural dynamics in the Urnfield period and the Middle Ages in the Southern Netherlands. editor / F. Theuws ; N.G.A.M. Roymans. Amsterdam, 1999. pp. 33-62 (Amsterdam archaeological studies; 4). 

Stijn Arnoldussen and Richard Jansen,Iron Age habitation patterns on the southern and northern Dutch Pleistoce coversand soils: The process of settlement nucleation, in: M. Meyer (eds.), Haus – Gehoft – Weiler – Dorf. Siedlungen der Vorromischen Eisenzeit im nordlichen Mitteleuropa, (Berliner Archaologische Forschungen 8), Berlin, 279-297, https://www.academia.edu/1091725/Arnoldussen_S_and_R_Jansen_2010_Iron_Age_habitation_patterns_on_the_southern_and_northern_Dutch_Pleistocene_coversand_soils_the_process_of_settlement_nucleation_in_M_Meyer_eds_Haus_Gehoft_Weiler_Dorf_Siedlungen_der_Vorromischen_Eisenzeit_im_nordlichen_Mitteleuropa_Berliner_Archaologische_Forschungen_8_Berlin_279_297

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De Mulder, Guy, and Jean Bourgeois, ‘Shifting Centres of Power and Changing Elite Symbolism in the Scheldt Fluvial Basin during the Late Bronze Age and the Iron Age’, in Tom Moore, and Xosê-Lois Armada (eds), Atlantic Europe in the First Millennium BC: Crossing the Divide (Oxford , 2012; online edn, Oxford Academic, 16 Mar. 2015), https://doi.org/10.1093/acprof:osobl/9780199567959.003.0013

Sørensen MLS, Rebay-Salisbury K. A Brief History of Urns, Urnfields, and Burial in the Urnfield Culture. In: Death and the Body in Bronze Age Europe: From Inhumation to Cremation. Cambridge University Press; 2023:15-35. https://www.cambridge.org/core/books/abs/death-and-the-body-in-bronze-age-europe/brief-history-of-urns-urnfields-and-burial-in-the-urnfield-culture/C971ED11A10E95B2E07F3E4C214F7015

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[74] Cremation involves incinerating the body to ash, inhumation is the burial of the body intact, and a tumulus is a burial mound that can contain an inhumation or cremation within it. The primary difference is the method of body disposal: cremation uses heat, inhumation uses burial, and a tumulus is a physical structure for either method. 

Burial TypeProcessRemainsTiming
CremationThe body is, incinerated, leaving behind bone fragments (ashes)Ashes are preserved in an urn, scattered, or buried.A service can happen anytime around the death, allowing for more flexible scheduling. 
InhumationThe body is buried intact in a cemetery or mausoleum.The body remains whole and is placed in a coffin or casket.Usually happens relatively quickly after death.
TumulusA burial mound, also called a barrow, is constructed over the deceased’s remains.A tumulus can house either an inhumation (buried intact) or the ashes from a cremation.It is an ancient structure and historical monument for commemorating and honoring the dead

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Fontijn, David & Fokkens, Harry. (2006). The emergence of Early Iron Age ‘chieftains’ graves’ in the southern Netherlands: reconsidering transformations in burial and depositional practices, Pages 354 – 373 in Colin Haselgrove and Rachel Pope, eds, The earlier Iron Age in Britain and the Near Continent, Oxford: Oxbow Books, 2007, https://www.researchgate.net/publication/333124942_The_emergence_of_Early_Iron_Age_’chieftains’_graves’_in_the_southern_Netherlands_reconsidering_transformations_in_burial_and_depositional_practices

Eugène Warmenbol, The Early Iron Age in Belgium, 201- 220, in Robert Schumann & Sasja van der Vaart-Verschoof, eds, Connecting Elites and Regions Perspectives on contacts, relations and differentiation during the Early Iron Age Hallstatt C period in Northwest and Central Europe, Leiden: Sidestone Press, 2017, https://www.sidestone.com/openaccess/9789088904424.pdf

Richard Jansen and Sasja van der Vaart-Verschoof, A cluster of chieftains’ graves in the Netherlands?, 127-144,  in Robert Schumann & Sasja van der Vaart-Verschoof, eds, Connecting Elites and Regions Perspectives on contacts, relations and differentiation during theEarly Iron Age Hallstatt C period in Northwest and Central Europe, Leiden: Sidestone Press, 2017, https://www.sidestone.com/openaccess/9789088904424.pdf

Van Beek, Roy and Arjan Louwen, Urnfields on the Move: Testing Burial Site-Settlement Relations in the Eastern Netherlands (c. 1100-500 BC), Archäologisches Korrespondenzblatt 42, 2012, 41-61, https://www.researchgate.net/publication/259801388_Beek_R_vanA_Louwen_2012_Urnfields_on_the_move_Testing_burial-site_settlement_relations_in_the_eastern_Netherlands_c_1100-500_BC_Archaologisches_Korrespondenzblatt_42-1_41-60

Roymans, N.G.A.M. ; Kortlang, F.P. / Urnfield symbolism, ancestors and the land in the Lower Rhine Region. Land and ancestors. Cultural dynamics in the Urnfield period and the Middle Ages in the Southern Netherlands. editor / F. Theuws ; N.G.A.M. Roymans. Amsterdam, 1999. pp. 33-62 (Amsterdam archaeological studies; 4). 

[79] Fernández-Götz, M. Urbanization in Iron Age Europe: Trajectories, Patterns, and Social Dynamics. J Archaeol Res 26, 117–162 (2018). https://doi.org/10.1007/s10814-017-9107-1 , https://www.pure.ed.ac.uk/ws/portalfiles/portal/42324611/Fernandez_Gotz_Urbanization_in_Iron_Age_Europe_Journal_of_Archaeological_Research.pdf

Halstatt Culture, Wikipedia, This page was last edited on 6 October 2025, https://en.wikipedia.org/wiki/Hallstatt_culture

[80] Fokkens H. The genesis of urnfields: economic crisis or ideological change? Antiquity. 1997;71(272):360-373. doi:10.1017/S0003598X00084970, https://www.researchgate.net/publication/28646976_The_genesis_of_urnfields_Economic_crisis_or_ideological_change

Heinrich G.H. Härke, Settlement Types and Settlement Patterns in the West Hallstatt Province: An Evaluation of evidence from Excavated Sites, BAR International Series 57 1979, https://www.academia.edu/477674/Settlement_types_and_settlement_patterns_in_the_West_Hallstatt_province_British_Archaeological_Reports_S57_Oxford_BAR_1979

Härke, Heinrich G. H.. Settlement Types and Settlement Patterns in the West Hallstatt Province: An Evaluation of Evidence from Excavated Sites. United Kingdom: British Archaeological Reports, 1979

Robert Schumann & Sasja van der Vaart-Verschoof, eds, Connecting Elites and Regions Perspectives on contacts, relations and differentiation during theEarly Iron Age Hallstatt C period in Northwest and Central Europe, Leiden: Sidestone Press, 2017, https://www.sidestone.com/openaccess/9789088904424.pdf

van der Vaart-Verschoof S, Schumann R. Connected by More Than Exceptional Imports: Performance and Identity in Hallstatt C/D Elite Burials of the Low Countries. European Journal of Archaeology. 2020;23(1):22-42. doi:10.1017/eaa.2019.44 , https://www.cambridge.org/core/journals/european-journal-of-archaeology/article/abs/connected-by-more-than-exceptional-imports-performance-and-identity-in-hallstatt-cd-elite-burials-of-the-low-countries/B30CD513E70AAC974511533326A07E12

Fernandez-Gotz, M 2018, ‘Urbanization in Iron Age Europe: Trajectories, patterns, and social dynamics’,

Journal of Archaeological Research, vol. 26, no. 2, pp. 117–162. https://doi.org/10.1007/s10814-017-9107-1

Looking at the Griff(is)(es)(ith) Y-DNA Phylogenetic Gap Associated with the Meuse and Rhine River Watershed from the Bronze Age Onward – Part Four

This fourth part of the story focuses on possible influences during and after the bronze age, up to approximately 650 CE, the estimated birth date of the most recent common ancestor that is asociated with haplogroup G-7648 at the end of the phylogenetic gap. Some of these influences are:

  • The enduring effects of the Bronze Age Bottleneck;
  • The enduring impact of the Corded Ware and Bell Beaker migrations on limiting the proliferation of G haplogroup subclades in susequent generations; and
  • Environmental impacts and changes in the delta landscape.

The 2,850 year Gap between G-FGC7516 and G-Z6748:  The most common recent ancestor associated with G-FGC7516 was born around 2200 BCE. The next genetic ancestor on the Griff(is)(es)(ith) YDNA line was associated with the genetic SNP mutation defining the G-Z6748 haplogroup, 2,850 years later. This gap of undocumented YDNA ancestors represents about 95 generations.

The Enduring Impact of YDNA Demographic Patterns Restricting G2a Subclade Proliferation in the Muese Rhine Watershed Area

The migratory path of the YDNA Griff(is)(es)(ith) lineage essentially followed the contours of the Danube and Rhine Rivers through time. The general migratory paths along these rivers roughly correspond with the two major phylogenetic gaps that have been discussed in six prior stories.

The lack of identified YDNA subclades documenting the migratory path in the Meuse Rhine watershed area can be partly attributed to three major demographic factors that limited the subsequent growth of G2a subclades, particularly the Griff(is)(es)ith) paternal line:

  1. the persistence and admixture of I2 and C1 hunter-gatherer haplogroups with G2a farming haplogroups in the late neolithic;
  2. the enduring impact of R1b Bell Beaker migrating groups into the area in the post Neolithic and early Bronze age; and
  3. the continued dominance of R1b social and cultural factors on G2a subclade growth.

1. Unique Persistence and Admixture of Hunter-Gatherer and Farming Y-Haplogroups

The Meuse-Rhine watershed area exhibited a distinctive demographic pattern compared to much of Europe when early G2a haplogroup farmer groups initially migrated into the area. There was a significant, long-term persistence of local hunter-gatherer YDNA ancestry. For thousands of years, the Rhine-Meuse region (covering the western and central Netherlands, Belgium, and western Germany) maintained a population with very high hunter-gatherer ancestry, up to fifty percent, much longer than surrounding areas, due to limited integration of early farmer ancestry (e.g. G2a lineages). The unique persistence and admixture of hunter-gatherer and farming Y-haplogroups is an unique characteristic of the Meuse-Rhine area throughout the Neolithic and Chalcolithic periods. [1]

The Y-DNA haplogroups representing the local mix of European hunter-gatherers and early European farmers were primarily I2C1, and G2a. These YDNA lineages reflect the blending of Mesolithic hunter-gatherer males and Neolithic agriculturalist males before the large influx of Steppe ancestry in the Bronze Age. The main I2 Y-DNA lineage was found among Western, Central, and Eastern European Mesolithic hunter-gatherers. They persisted widely through the Neolithic and into the Copper Age, especially in areas with continued high hunter-gatherer ancestry. Haplogroup C1 was much rarer haplogroup but was detected in some Western and Northern European Mesolithic and Early Neolithic remains, showing deep Paleolithic roots in Europe. [2]

The G2a Early European Farmer (EEF) male lineage, the lineage representing the Griff(is)(es)(ith) paternal line, was among the Anatolian/Levantine-derived early Neolithic farmers who spread agriculture into Europe through northern and sourthern Europe, especially in Linearbandkeramik (LBK), Cardial, and other early Neolithic cultures. [3] The F and J haplogroups, less common but also observed among some early farmer groups, were present particularly in southern and southeastern Europe. [4]

Source: Andrew Zeilstra and Johanna Knop, Heightened Interaction Between Neolithic Migrants and Hunter-Gatherers in Western Europe, 29 May 2020 Press Release, Max Planck Institute of Geoanthropology, https://www.shh.mpg.de/1713184/haak-french-dna#_ftnref4

As mentioned in the video above, regions such as the Meuse-Rhine area, Central Europe, and France showed intermixed communities during the Neolithic and Chalcolithic periods, with both I2 (hunter-gatherer) and G2a/F (farmer) haplogroups detected among males. The presence of both I2 and G2a haplogroups in later Neolithic era sites indicates communities with significant admixture between local hunter-gatherers and arriving farming peoples.

In regions with high western hunter-gatherer and early European farmer admixture, especially before the steppe-related R1b spread, Y-DNA lineages I2 and G2a were typical representatives of the local male genetic landscape. As reflected in table one, ancient DNA (aDNA) from the Neolithic and Chalcolithic Meuse-Rhine region shows that males predominantly belonged to haplogroups I2aR1b-V88, and C1a—all linked to European hunter-gatherers [5]

Table One: Haplogroup Presence in the Mesue Rhine Watershed Area

Period/PopulationY-DNA HaplogroupsDescription
MesolithicI2a, C1a, R1b-V88Western Hunter-gatherer (WHG) dominant
Early Neolithic MixI2a, C1a, R1b-V88 + minor representation of G2a40–50% WHG, 50–60% Early Eurpean Farmer (EEF) admixture
Post-Neolithic BeakerR1b-L151Steppe ancestry dominant, local mix wanes

The G2a haplogroup was present in the local Neolithic and Chalcolithic populations of the Meuse-Rhine area, but it was not as dominant there as in many other parts of Europe. While early European farmers across the continent are strongly associated with G2a, ancient DNA from the Rhine-Meuse region during the Neolithic instead shows a much higher persistence of hunter-gatherer lineages such as I2a and C1a, with G2a present but at relatively lower frequency. This may partly explain the dearth of discovered subclades in the Griff(is)(es)(ith) paternal migratory line of descent through this area of western Europe.

The great winner during the Neolithic period was haplogroup I2a, which consistently shows up alongside G2a in most Neolithic sites tested to date (Starčevo, Körös, Lengyel, LBK, Cardium Pottery, Megalithic), and seem to increase in frequency over time and as one moves towards Northwest Europe.

“(A)lthough I2a was just one of many Mesolithic hunter-gatherers’ lineages in Europe when agriculturists arrived, it is the only one that readily embraced the new lifestyle and managed to supersede the original farmers in number. I2a’s destiny was not only linked to its ability to chum with G2a, but we could say that G2a farmers catalysed I2a’s success. I2a people integrated G2a tribes, learned the new Neolithic techniques from them and became so good at them that over time the student overtook the master. [6]

The Rhine-Meuse region’s river-dominated landscapes significantly shaped the adoption and adaptation of Neolithic farming practices through ecological constraints, specialized subsistence strategies, and the blending of hunter-gatherer and farming cultural practices. The dynamic wetland/riverine environment (marshes, peat bogs, and seasonal floods) hindered full-scale Neolithic agriculture. Communities developed a mixed subsistence strategy combining:

  • Limited crop cultivation on elevated river dunes/levees;
  • Cattle husbandry optimized for wetland conditions (grazing on salt marshes, occasional winter foddering); and
  • Persistent hunting, fishing, and foraging in resource-rich aquatic ecosystems.

This “extended broad spectrum” approach allowed populations to exploit the landscape without abandoning traditional Mesolithic practices. [7]

2. Impact of Bell Beaker Culture Migration and YDNA Replacement: A Local Admixture Event Resulting in New Populations with Dominant Steppe Ancestry

During the arrival of the Corded Ware complex, local individuals in this region adopted elements of the material culture but exhibited very little steppe ancestry, unlike Corded Ware sites elsewhere. Bell Beaker-associated populations in this region around 2500 BCE were formed by a mixture of Corded Ware-related migrants with steppe ancestry and the persistent local ‘Neolithic substrate’, with the local contribution modelled at 9 to 17 percent. Bell Beaker men in the Rhine-Meuse region mostly carried the R1b-L151 (especially P312) Y-chromosome, absent in earlier Neolithic populations but present among Central European Corded Ware groups, indicating a strong but not exclusive external influx. [8]

In much of Northwestern Europe, Bell Beaker and R1b haplogroup expansion involved almost total population and male lineage replacement. In contrast, as reflected in table two, the Rhine-Meuse area saw a transformative but partly local admixture event, resulting in new populations with dominant steppe ancestry (about 83–91 percent) but still a recognizable input from the enduring local hunter-gatherer- early European farmer influenced population. This “fusion zone” became a launching pad for further population expansions into regions such as Britain, where the Bell Beaker-associated transformation was even more complete. [9]

Table Two: Distinctive Demographic Impact in Muese Rhine Watershed Area

RegionPre-Beaker PopulationBeaker/Early Bronze Age PatternEstimated Ancestry TurnoverLocal Genetic Input
Meuse-RhineHunter-Gatherer ‘Rich Mix’ with Early European FarmersBell Beaker: R1b, steppe ancestry + local admixture83–91% turnover9–17%
BritainNeolithic FarmersNear-total replacement by R1b-rich, steppe Beaker90–100% turnover0–9%
Central EuropeNeolithic Farmers + WHGR1b predominance, steppe ancestry via Corded WareVery highMinimal

The Meuse-Rhine region stands out for its partially blended transformation during the Beaker phenomenon, marked by fusion rather than just replacement, with steppe-derived R1b Y-haplogroups and ancestry still predominating in the end. This admixture pattern was unusual for Europe. In most regions, Neolithic migration led to the overwhelming dominance of farmer-origin lineages (e.g. G2a), with much lower persistence of I2 or other hunter-gatherer Y-DNA. In the Meuse-Rhine, admixture continued for centuries, and the local hunter-gatherer Y-haplogroups persisted at high levels until the arrival of Bell Beaker people with predominantly steppe and R1b-L151 ancestry.

3. The Enduring Impact of Post-Bronze Age Influences

The decline of G2a subclades in the Netherlands was not a single event but a cumulative process over generations. It began with the Bronze Age migrations that replaced many Neolithic paternal lineages, and was further influenced by demographic, environmental, and social changes in subsequent time periods.

Pre-existing populations, including those with high frequencies of G2a, may have faced bottlenecks, diseases, or environmental changes that reduced their numbers, making them more susceptible to genetic replacement. For example, archaeological evidence from the Netherlands points to periods of population decline and settlement abandonment during the Roman Empire’s collapse, which further complicated and reshaped the genetic landscape. [10]

The Impact of Patrilineal and Patrilocal (Virolocal) Social Structures on YDNA Diversity

Many of the Bronze age and cultures in subsequent achealological periods were patrilineal and patrilocal. Patrilineal and patrilocal (virolocal) social structures leave distinct, observable signatures on human Y-chromosome diversity.  Patrilineal and patrilocal/virolocal systems lead to reduced Y-chromosome diversity within groups and lineages, with distinct genetic clustering matching societal divisions.

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In the Netherlands specifically, studies suggest that much of the change in Y-chromosome frequencies over the following centuries was due to genetic drift and stable patrilocal social structures that reinforced the dominance of R1b haplogroups in their communities over generations. This led to the persistence and reinforcement of newer lineages, rather than regular influx or replacement, so that the effect of the R haplogroup steppe expansion was locked in for centuries. [11]

Genetic drift, the random fluctuation of gene variant frequencies in a population, would have also contributed to the decline of G2a. In small, isolated communities, the loss of certain lineages due to chance events or the failure of a male line to reproduce can have a significant impact. Shifts in reproductive dynamics, including new social structures and marriage patterns brought by the migrating populations, may have further disadvantaged the older G2a paternal lines. [12]

Subsequent migrations during the Iron Age and the early Middle Ages—such as the arrival of Germanic tribes (including Angles, Saxons, and Franks)—continued to shape the genetic makeup of the Low Countries in the Muese Rhine watershed area. This further diluted or replaced the genetic signatures of earlier groups, cementing the marginalization of older lineages like G2a[13]

A recent study by Eveline Altena and associates provides an in-depth look at paternal genetic continuity in the Netherlands across a span of 1,300 years. The key findings indicate remarkable stability of male lineages. The team analyzed the Y-chromosomes of 348 men from 13 Dutch locations dated 500 CE–1850 CE, alongside modern YDNA data, to trace paternal ancestry across millennia, see illustration one. [14]

Illustration One: Observed Y-Haplogroup Frequencies in Three Historical Periods in the Middle Ages in the Netherlands

Click for Larger View | Source: Figure 3 in Altena, Eveline, et al., The dutch Y-chromosome from the early middle ages to present day. Archaeol Anthropol Sci 17, 116 (2025). https://doi.org/10.1007/s12520-025-02224-4

The study found that the male population of the Netherlands showed limited change from the Early Middle Ages to modern times. Based on the study’s findings, fluctuations in haplogroup frequencies mostly resulted from genetic drift rather than large-scale migration or population replacement.

The study also supports the impression that the medieval Netherlands practiced patrilocality—men stayed put in localized areas and women moved for marriage. Regional male lineages were reinforced and preserved by limited male mobility, while mitochondrial (female) variation was more diffuse.

The population substructure and gradients for many of the individual YHGs (Y haplogroups) we found in our study are in strong contrast with the apparent lack of genetic-geographic patterns for mtDNA data . . . . This could be an indication of different demographic histories for women and men. One could think, for example, of the patrilocal residence system, which is typical for farming societies, such as the Dutch. In these societies sons stay with their family and daughters move to the residence of their husbands. Also, genetic drift may have acted differently on mt-DNA than on Y-chromosomes.” [15]

Regional male lineages were reinforced and preserved by limited male mobility, while mitochondrial (female) variation was more diffuse. Over centuries, Dutch male lineages were shaped less by mass migration than by chance and social practice, setting them apart from regions with more turbulent demographic histories. [16]

In an earlier published article in 2019 and 2020, Altena and associates completed a geographic analysis of Y-chromosome haplogroup (YHG) distribution across the Netherlands. Using data from 2,085 males and integrating information from northern Belgium, the study found distinct geographic patterns in Y-chromosome distribution, with multiple Y-haplogroups showing significant clinal frequency gradients (i.e., gradual changes in frequency across regions). [17]

While previous research found limited or no mitochondrial DNA (maternal-line) spatial patterns or substructure, the pronounced Y-chromosomal substructure points to different population histories for men and women in the Netherlands. Contrasting male and female lineage patterns suggest sex-biased demographic histories, with male lineages undergoing more pronounced geographic differentiation, perhaps reflecting historical migration, social, or cultural practices.

Prediction surface maps were used to visualize the complex distributions of individual Y-haplogroups in detail, revealing non-random patterns throughout the country for almost every haplogroup examined. Y-chromosome diversity in the Netherlands shows a significant micro-geographic structure, with several haplogroups (e.g., R1b variants) displaying regional gradients.

Illustration Two: Prediction Surface Maps of the Four Most Frequent Y Haplogroups in the Dutch Dataset

Click for Larger View | Source: Figure 5,Prediction surface maps of the four most frequent (sub-)YHGs in the Dutch dataset in phylogenetic order, inAltena E, Smeding R, van der Gaag KJ, Larmuseau MHD, Decorte R, Lao O, Kayser M, Kraaijenbrink T, de Knijff P. The Dutch Y-chromosomal landscape. Eur J Hum Genet. 2020 Mar;28(3):287-299. doi:10.1038/s41431-019-0496-0 . Epub 2019 Sep 5. Erratum in: Eur J Hum Genet. 2020 Mar;28(3):399. https://doi.org/10.1038/s41431-019-0496-0

Y-haplogroups “G-M201, J2-M172, R1b-M269, and R1b-S116 increase from north to south, R1b-M405 Total and R1b-L48 increase from south to north, I-M170 increases from southwest to northeast and R1b-S116 Total, R1b-U152 and R1b-M529 increase from northeast to southwest. . . . From all the YHGs for which prediction surface maps were created, only YHG R1b-M529 is more or less evenly distributed over the Netherlands. All other YHGs show more distinct patterns of distribution.” [18]

Altena and associate researchers point out that the haplogrup G-M201 which the Griff(is)(es)(ith) paternal line is part of “is relatively rare in Europe, with average proportions below 5% in northwestern Europe, which is consistent with our findings. Because proportions are low throughout the most of Europe, there is no clear gradient, but overall it increases from northwest to southeast (in the Netherlands and Belgium) . . . .“. [19]

Today’s clear-cut geographic patterns in Dutch Y-DNA formed late, likely as the result of more recent events and not from deep medieval roots. [20]

The Use of Archaeological Time Periods Associated with this Phylogenetic Gap

The loose reference at the begining of this story to a time period “during and after the bronze up to 650 CE” can be referenced by and viewed through the following archaeological time periods in table three. The approximate dates for the three Bronze Age periods, the two periods of the Iron Age, the Roman era and the Merovian perod in the Meuse-Rhine river watershed are based on archaeological and historical studies. These approximate time ranges reflect scholarly consensus for distinctive time periods for the Meuse-Rhine watershed, though there may be slight regional variation within the area.

Utilizing archaeological time periods offers the advantage of organizing human history into discernible phases. Archaeological time periods (like the Stone, Bronze, and Iron Ages) help organize the development of human societies in a clear chronological framework, making it easier to compare changes across time and regions. These periods enable archaeologists to interpret artifacts and features within a broader cultural and technological context, offering insight into the evolution of societies, technologies, and economies. Standardized periods facilitate the communication of archaeological findings to both academic communities and the public, enhancing understanding of complex historical developments. [21]

However, there are obvious limitations associated with the use of archaeological time periods. Broad periods like “Stone Age” or “Iron Age” can mask local and regional diversity, as technological advancements and cultural practices often occurred at different times in different places, leading to overlaps that the periods cannot accurately reflect. Assigning boundaries to these periods involves subjective decisions based on material culture, which can introduce bias and debate among scholars. [22]

Table Three: Archaeological Periods in the Meuse Rhine Watershed Area During the Phylogentic Gap for the Griff(is)(es)(ith YDNA Lineage

Archaeological PeriodApproximate DatesArcheaological Description
Early Bronze Age2200–1600 BCE At the beginning of the Bronze Age, the Meuse-Rhine area was at a cultural crossroads and was influenced by broader European traditions. The Beaker culture dominated much of Western and Central Europe and is considered a transitional culture between the late Neolithic and the early Bronze Age. While primarily present further east, the Corded Ware culture also influenced the early Bronze Age in parts of the region, with archaeologists sometimes tracing burial rituals back to these roots. 

Communities were settled along elevated stream ridges and river banks, engaging in mixed farming and livestock herding, reflecting the broader transition to more sedentary life seen kin other parts of north and central Europe.

There was the persistence of distinctive local cultures with high levels of hunter-gatherer ancestry, in contrast to the rest of Europe where farming ancestry became dominant earlier. The area’s wetlands and rivers promoted more isolated populations with limited early integration of outside groups. [23]
Middle Bronze Age1600–1300 BCE The region experienced a significant increase in settlements, with numerous house sites discovered in the river delta, facilitated by technological advances such as improved bronze tools and animal husbandry.

Elp culture (northern sector): Located in the northern and eastern Netherlands, this culture existed from approximately 1800 to 800 BCE. Hilversum culture (southern sector): Centered in the southern Netherlands and northern Belgium, this culture has been linked to the Wessex culture of southern England.

Long-distance contacts and cultural exchange increased, reflected in the archaeological record by the emergence of ‘pan-European’ material culture, but settlements declined toward the end of this period hints at possible floods or ecological stresses.

Large-scale deforestation for agriculture is evident, paralleling the broader trend of expanding farmland and increased social complexity. [24]
Late Bronze Age1300–800 BCE The Urnfield culture became the dominant cultural tradition across Central Europe and extended into the Meuse-Rhine area during the Late Bronze Age. A local sub-group known as the Lower Rhine Group developed within the Urnfield tradition. The Urnfield period saw an increase in social stratification and widespread trade networks. It is also characterized by the appearance of fortified hilltop settlements and advanced bronze metalworking.

A sharp decline in settlements is noted in the region, partly attributed to climate change, flooding, and changes in river dynamics, but also possibly to changing social structures or shifts toward more mobile ways of life.

Evidence points to sustained but reduced habitation, with human activity focused on certain elevated or defendable sites. This aligns with the general Bronze Age trend toward urbanization and sociopolitical stratification, but environmental instability made large, permanent settlements less sustainable.

There was significant contact with neighboring areas, visible in material culture and genetics, corresponding with the general increase in social complexity and long-distance exchange in the wider Bronze Age world. [25]
Iron Age [26]Early Iron Age (Hallstatt period): 800-500 BCE

Late Iron Age (La Tène period): 450 -50 BCE
The Meuse-Rhine area served as a transitional zone between the Celtic south (La Tène) and the Germanic north (Hallstatt), resulting in a unique blend of traditions. The area saw continuity from prehistoric populations with new influences entering gradually, rather than abrupt cultural shifts. Settlements in the Meuse-Rhine Iron Age area were typically small and lacked the major fortified oppida found further south in La Tène regions.

Archaeological surveys note a decline or transformation in settlement patterns in the early Iron Age, with fewer large settlements and more dispersed smaller sites, contrasting with the denser occupation of the Late Bronze Age. [27]

The region experienced significant environmental changes such as increased flooding and sedimentation in the river valleys, likely due in part to enhanced human land use such as farming and deforestation.

Archaeological evidence shows locally produced material goods, e.g. pottery, basic iron tools, and simple burial rituals rather than elaborate grave goods. The adoption of La Tène cultural elements—occurred only to a limited degree, reflecting modest trade and cultural interaction with Celtic areas.
Roman Era [28] 55 BCE–459/461 CE The Roman Era in the Meuse-Rhine Watershed was defined by advances in water management, the creation of complex settlements along the Roman frontier, multicultural dynamics, and a blend of indigenous and Romanized landscape use, leaving a distinctive archaeological legacy.
Late Antiquity/Post-Roman Transition460–500 CECollapse of Roman state authority, significant depopulation, urban and rural structures deteriorated, the Roman limes (frontier) ceased to function [29]
Early Merovingian Period500–650 CERegion became part of Frankish kingdom, under Merovingian rule. Settlement patterns shifted, with new rural settlements developing—sometimes continuing from or near former Roman sites—and local communities reorganizing around new Frankish leadership structures.

The area retained the Roman “pagi” (district) administrative structure, following both old Roman and Germanic tribal traditions in leadership and justice, with leaders initially selected by local warriors.

The chaotic social context, continually punctuated by small-scale warfare, fostered the rise of a local warrior elite—the precursors to later medieval nobility—rewarded with land for military service. [30]

While archaeological time periods help bring order and clarity to the study of human prehistory and history, they are best used as flexible frameworks rather than rigid chronological boundaries, always considering the limitations and regional variations they entail. Regional chronologies—such as division into Early, Middle, and Late Bronze Age—are not uniform and may overlap or differ in exact dating due to differing archaeological traditions and the tempo of cultural change and adoption of technicalogcal change as reflected in archaeological artifacts.

The limitations of utilizing ‘time periods’ can be specifically demonstrated when looking at the ‘bronze age’ in the Meuse-Rhine watershd area. The Bronze Age in Europe shows significant regional variation in both its starting and ending dates, as well as in developmental phases. The earliest Bronze Age developments in Europe occurred in the Aegean and southeast Europe, spreading gradually northwest and north. Central and northern regions (e.g. Scandinavia and the British Isles) entered the Bronze Age later due to environmental, technological, and cultural factors.

Archaeologist Harry Fokkens has called for a reassessment of the criteria used to define archaeological periods, advocating for a more multi-faceted approach that combines absolute carbon dating with meaningful archaeological markers, such as changes in settlement organization, technology, and social structure. He urges, as summarized in table two, archaeologists to move beyond simply updating dates and to critically reflect on the cultural significance of period boundaries (see table four). [31]

Table Four: Fokkens’ Recommendations for Sustantiation of Archeological Periods

AspectCurrent PracticeFokkens’ Proposed Revision
Primary EvidenceBurial data and bronzesInclusion of Settlement patterns, houses, material culture
Theoretical BasisMigration/culture-change modelsLocalized cultural continuity analysis
Chronological ToolsRadiocarbon datingCombined: carbon dating & cultural/economic shifts
TransparencyAssumptions rarely scrutinizedExplicit criteria and peer-reviewed frameworks

Fokkens argues that the division of the Bronze Age into different cultures and phases was predominantly based on burial data, the nature and use of pottery, and migration theories and culture-change models that often relied on weak evidence for social, religious, or spiritual changes, which, while detailed, lacked scientific credibility. Other cultural phenomena like settlement patterns and house types were underutilized or inconsistently considered. He argues that the bronze age in the Netherlands existed roughly between 1800 – 800 BCE.

” … (W)e should move the line dividing the Neolithic and Bronze Age towards 1800 BC. This is the absolute end of the Beaker traditions and the start of entirely new ones, maybe not so much in a technological sense but in social, economical and ritual aspects of culture. I therefore propose to indicate the period between 2900 and 1800 BC as the Late Neolithic, the period of the Beaker Cultures. Late Neolithic A is the period of the Single Grave Culture including AOO Beakers (2900-2500 BC), Late Neolithic B the period of the Bell Beaker Culture (2500- 2000), Late Neolithic C the period of the Barbed Wire Beaker Culture (2000-1800).

The Early Bronze Age, as I see it, is marked by the development of several traditions that differ from the Late Neolithic practices sufficiently to suggest that a fundamental change in several dimensions of culture occurred simultaneously. Housing traditions changed and possibly associated economic traditions, burial traditions, deposition practices and pottery traditions. At the same time new traditions start which continue for the next 700 years.” [32]

While Fokkens does not propose a specific alternative chronological framework with fixed phases and dates for the Dutch Bronze Age, the chronological boundaries of the Bronze Age in the Netherlands are generally considered to span from around 2000 BCE to 800 BCE based on research studies and archaeological periodization. While some sources mention minor earlier traces or slight local variation, most scholarly consensus places the start at about 2000 BCE, marked by the first appearance of bronze objects and the Barbed-Wire Beaker culture, and the end at around 800 BCE, when the transition to the Iron Age occurs.

Martin Furholt, makes a similar agument when discussing the European Neolithic Period. He highlights how recent advances in archaeogenetics—the analysis of ancient DNA—have revolutionized the understanding of mobility and social dynamics in Neolithic Europe. He also critically examines the interpretations and intellectual frameworks that have emerged from this new data. [33]

Furholt argues that the enthusiasm for genetic data may lead to simplistic, macro-level migration narratives (e.g., mass migrations or population replacements), reminiscent of earlier, now outdated models from the early 20th century that treated archaeological cultures as homogenous, biologically-defined groups. He stresses the importance of integrating genetic findings with archaeological and anthropological theory, warning against equating genetic patterns directly with social identity or assuming that genetic groups correspond neatly to archaeological cultures.

Furholt recommends moving beyond the idea of migration as a simple mass movement and instead encourages nuanced, case-by-case studies that combine diverse lines of evidence, such as material culture, burial practices, and genetic data. Furholt calls for better interdisciplinary collaboration between archaeologists and geneticists so that social, cultural, and biological data can together produce more sophisticated and accurate models of Neolithic social change. He calls for better interdisciplinary collaboration between archaeologists and geneticists so that social, cultural, and biological data can together produce more sophisticated and accurate models of Neolithic social change.

Bronze Age Cultures in the Muese-Rhine Watershed

The Bell Beaker phenomenon was never a monolithic culture but rather a complex network of related groups with substantial regional variation. As Bronze Age societies developed, these regional groups either merged with or were absorbed by neighboring cultures, leading to the fragmentation and eventual disappearance of distinct Bell Beaker identities. Key cultures marking the start and end of the European Bronze Age differ by region but show several clear patterns. [34]

The exact cause of the decline of the Bell Beaker culture is a subject of ongoing debate among archaeologists and historians, and there are various theories proposed. It is generally understood that the Bell Beaker culture, which flourished in Europe from around 2800 to 1600 BCE, eventually faded and was succeeded by new cultural norms and the onset of the Bronze Age. [35]

The Bell Beaker culture did not vanish abruptly. It was succeeded by emerging Bronze Age cultures such as the Únětice culture in Central Europe and various regional Bronze Age societies elsewhere. This transition often involved the blending of Bell Beaker traditions with those of incoming or neighboring groups, leading to new cultural identities and practices.

The major cultural groups succeeding the Bell Beakers in the Meuse-Rhine watershed were the Hilversum Culture during the Early Bronze Age, followed by the Urnfield Culture in the Middle to Late Bronze Age, leading into the various Early Iron Age societies. As the Urnfield Culture transitions to the Early Iron Age, archaeological evidence suggests further shifts, possibly involving early Hallstatt influences and precursor groups to the Celts (see table five). It is likely that the roughly 95 undocumented generations of the Griff(is)(es)(ith) paternal line were part of many of the cultures listed in table five.

Table Five: Major Cultural Groups in the Meuse Rhine Watershed Area by Archaeological Period

CulturePeriod (BCE)Archaeological Period and Location
Corded Ware~ 2900–2400Late Neolithic/Early Bronze Age
Bell Beaker~ 2500–2100Early Bronze Age; mix of local/steppe ancestry
Barbed Wire Beaker~ 2000–1800Early Bronze Age – Late Bell Beaker variant in region
Hilversum~ 1870–1050Middle Bronze Age, south Netherlands/Belgium
Elp~ 1800–800Middle to Late Bronze Age, north/east Netherlands
Tumulus~ 1600–1200Middle Bronze Age, tumulus (barrow) burials
Lower Rhine Urnfield~ 1300–750Late Bronze Age, cremation fields
Hallstatt~ 800–450 Early Iron Age (C/D proper); outgrowth of Urnfield, proto-Celtic
La Tène, Germani cisrhenani~ 450 – 50 Late Iron Age
Batavi, Cananefates, Tungri, Romanized locals~ 50 BCE – 400 CERoman Era
Frankish (merovingian), Saxons, Frisians~ 400 – 800 CEEarly Medieval

These cultural transformations set the stage for later groups in the Iron age such as the Nordwestblock, theorized as a distinct cultural area between Celtic and Germanic zones in the Lower Rhine and Meuse area. [36]

The Bell Beaker cultures (2700–2100) locally developed into the Bronze Age Barbed-Wire Beaker culture (2100–1800). In the second millennium BC, the region was the boundary between the Atlantic and Nordic horizons and was split into a northern and a southern region, roughly divided by the course of the Rhine. [37]

The Rhine River would continue to have an ecological impact on competing cultural and social groups and have an effect on the YDNA composition through susequent archaeological time periods. It would also have an impact on the absence of documented YDNA subclades associated with the migratory path of the Griff(is)(es)(ith) paternal line.

Parting Comments

The contested Meuse and Rhine River watershed area shaped and restricted the growth of the Y-chromosomal DNA phylogenetic tree of the Griff(is)(es)(ith) paternal line through dynamic interactions between ecological barriers and other social groups. Hunter gatherer groups, male-biased migrations, demographic impacts on subclade proliferation and geopolitical conflicts in each of the archaeological times periods.

The region’s wetlands and riverine landscapes resisted early Neolithic farming, preserving I2a and C1a Y-haplogroups linked to Mesolithic hunter-gatherers and limiting subclade proliferation of G2a haplogroups. Genetic continuity persisted until about 2500 BCE, , longer than in continental Europe, due to limited integration of Anatolian farmer females into local communities. This ecological resistance created a “genetic refugium” for older paternal lineages.

Despite adopting Corded Ware cultural practices, lowland Rhine-Meuse populations retained minimal steppe ancestry. However, Corded Ware associated R1a-M417 Y-haplogroups were introduced through male-dominated migrations, while retaining about 50 percent of local Neolithic ancestry. Limited female gene flow from steppe populations preserved high hunter-gatherer autosomal ancestry.

The formation of Bell Beaker groups through the mixing of local Rhine-Meuse populations (about 9–17%) and Corded Ware migrants led to the near complete Y lineage replacement of G2a YDNA groups. Bell Beaker associated males carried R1b-L151 subtypes (e.g., P312U106), replacing earlier Neolithic lineages such as the Griff(is)(es)(ith) YDNA line. Steppe-ancestry males again mixed with local females.

These patterns demonstrate how the Rhine-Meuse watershed’s contested history created a stratified YDNA phylogeny of dominant and minorty subclades, with clines and subclades reflecting a millennia of asymmetric gene flow, male-driven expansions, and ecological resilience.

An highly contested geographical region like the Meuse-Rhine watershed, coupled with its ecological influences, profoundly shaped phylogenetic YDNA trees over time, creating a genetic landscape marked by strong lineage turnover and historical layering. The unique combination of ecological complexity and frequent territorial disputes in this area led to several characteristic outcomes:

Persistent Ancient Minority Lineages Through Isolation

Swampy lowlands, river deltas, and ecological bottlenecks preserved ancient hunter-gatherer Y-haplogroups (such as I2a and C1a) for thousands of years longer than in neighboring regions. These ‘refugia effect zones’ resisted incoming male lineages from early farming and steppe cultures, leading to deep-rooted branches on local YDNA trees that survived alongside more recent YDNA subclades.

Abrupt Lineage Replacements via Male-Dominated Migrations

Periods of intense population turnover—such as the Corded Ware and Bell Beaker expansions—produced abrupt, near-complete Y-chromosome replacements marked by new dominant haplogroups like R1b-L151 and R1a-M417 in the archaeological record. These replacements rarely erased all earlier diversity, but pruned YDNA trees of specific Neolithic branches while fostering ‘star-like phylogenies’ indicative of founder effects and rapid expansions. [38]

Genetic Subclades Shaped by Political Borders

Long-standing borders acted as semi-permeable filters that constrained male gene flow. These boundaries, such as the Rhine and Meuse Rivers, imprint phylogenetic trees with localized subclades and long phylogentic trees with few branches, revealing the enduring effect of contested territories.

Frequent social and political fragmentation after various groups changed or collapsed magnified genetic drift and lineage bottlenecks in isolated settlements. This created micro-regional YDNA substructures, amplifyed minor lineages through stochastic processes while reducing overall haplogroup diversity in places with population contractions.

Accumulation of Genetic Layers in the ‘Long Duration’ of YDNA Genetic Time

As social groups vied for control over time, the region’s YDNA composition mirrored the area’s archaeological and historical complexity: every territorial upheaval left its signature as a branch or cluster on the YDNA phylogeny that may persist or be overwritten by subsequent events.

In combination, these impacts mean highly contested regions like the Meuse-Rhine watershed display phylogenetic YDNA trees marked by both ancient depth and recent star-phylogeny expansions, strong subregional differentiation, and evidence of recurrent lineage pruning and replacement driven by both ecological and sociopolitical forces.

The final part of this story will discuss the major cultural groups associated with the migratory path for the Griff(is)(es)(ith) paternal line succeeding the Bell Beakers through the Bronze Age, Iron Age, the Roman Era and the early medieval age.

Source:

Feature Banner: The banner at the top of the story features a map of the phylogenetic gaps discussed in the story. The maps was generated by taking a snapshop from the FamilyTreeDNA GlobetrekkerTM video of the migratory path of my YDNA descendants over time. The map shows the migratory path of selected most common recent ancestors and their respective estimated dates of birth. Another map in the banner depicts the Muesse and Rhine River watershed that is associated with this phylogenetic gap. . In addition, various cultures and features associated with time periods within this period of time are depicted.

[1] Iñigo Olalde, Eveline Altena, Quentin Bourgeois, Harry Fokkens, Luc Amkreutz, Marie France Deguilloux, Alessandro Fichera, Damien Flas, Francesca Gandini, Jan F. Kegler, Lisette M. Kootker, Kirsten Leijnse, Leendert Louwe Kooijmans, Roel Lauwerier, Rebecca Miller, Helle Molthof, Pierre Noiret, Daan C. M. Raemaekers, Maïté Rivollat, Liesbeth Smits, John R. Stewart, Theoten Anscher , Michel Toussaint, Kim Callan, Olivia Cheronet, Trudi Frost, Lora Iliev, Matthew Mah, Adam Micco, Jonas Oppenheimer, IrisPatterson, Lijun Qiu, Gregory Soos, J. Noah Workman, Ceiridwen J. Edwards, Losif Lazaridis, Swapan Mallick, Nick Patterson, Nadin Rohland, Martin B. Richards, Ron Pinhasi, Wolfgang Haak, Maria Pala, David Reich, Long-term hunter-gatherer continuity in the Rhine Meuse region was disrupted by local formation of expansive Bell Beaker groups, bioRxiv 2025.03.24.644985; doi: https://doi.org/10.1101/2025.03.24.644985, https://www.biorxiv.org/content/10.1101/2025.03.24.644985v1.full; also found at: https://pubmed.ncbi.nlm.nih.gov/40196638

[2] Szécsényi-Nagy Anna, Guido Brandt, Wolfgang Haak, Victoria Keerl, János Jakucs , Sabine Möller-Rieker, Kitti Köhler, Balász Gusztáv Mende, Krisztián Oross , Tibor Marton, Anett Osztás, Viktória Kiss, Marc Fecher, Gyögy Pálfi, Erika Molnár, Katalin Sebők, András Czene, Tibor Paluch, Mario Šlaus, Mario Novak, Nives Pećina-Šlaus, Brigitta Ősz, VandaVoicsek, Krisztina Somogyi, Gábor Tóth, Bernd Kromer, Eszter Bánffy, Kurt W. Alt. Tracing the genetic origin of Europe’s first farmers reveals insights into their social organization. Proc Biol Sci. 2015 Apr 22;282(1805):20150339. doi: 10.1098/rspb.2015.0339. PMID: 25808890; PMCID: PMC4389623 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC4389623/

António Faustino Carvalho, Eva Fernández-Domínguez, Eduardo Arroyo-Pardo, Catherine Robinson, João Luís Cardoso, João Zilhão, Mário Varela Gomes, Hunter-gatherer genetic persistence at the onset of megalithism in western Iberia: New mitochondrial evidence from Mesolithic and Neolithic necropolises in central-southern Portugal, Quaternary International, Volumes 677–678, 2023, Pages 111-120, ISSN 1040-6182,
https://doi.org/10.1016/j.quaint.2023.03.015 .
(https://www.sciencedirect.com/science/article/pii/S104061822300099X )

Primorac D, Šarac J, Havaš Auguštin D, Novokmet N, Bego T, Pinhasi R, Šlaus M, Novak M, Marjanović D. Y Chromosome Story-Ancient Genetic Data as a Supplementary Tool for the Analysis of Modern Croatian Genetic Pool. Genes (Basel). 2024 Jun 6;15(6):748. doi: 10.3390/genes15060748. PMID: 38927684; PMCID: PMC11202852 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11202852/

Western hunter-gatherer, Wikipedia, This page was last edited on 18 August 2025, https://en.wikipedia.org/wiki/Western_hunter-gatherer

Posth, C., Yu, H., Ghalichi, A. et al. Palaeogenomics of Upper Palaeolithic to Neolithic European hunter-gatherers. Nature 615, 117–126 (2023). https://doi.org/10.1038/s41586-023-05726-0

[3] Primorac D, Šarac J, Havaš Auguštin D, Novokmet N, Bego T, Pinhasi R, Šlaus M, Novak M, Marjanović D. Y Chromosome Story-Ancient Genetic Data as a Supplementary Tool for the Analysis of Modern Croatian Genetic Pool. Genes (Basel). 2024 Jun 6;15(6):748. doi: 10.3390/genes15060748. PMID: 38927684; PMCID: PMC11202852. (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11202852/

Szécsényi-Nagy Anna, et al. Tracing the genetic origin of Europe’s first farmers reveals insights into their social organization. Proc Biol Sci. 2015 Apr 22;282(1805):20150339. doi: 10.1098/rspb.2015.0339. PMID: 25808890; PMCID: PMC4389623 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC4389623/

Z. Hofmanová, S. Kreutzer, G. Hellenthal, C. Sell, Y. Diekmann, D. Díez-del-Molino, L. van Dorp, S. López, A. Kousathanas, V. Link, K. Kirsanow, L.M. Cassidy, R. Martiniano, M. Strobel, A. Scheu, K. Kotsakis, P. Halstead, S. Triantaphyllou, N. Kyparissi-Apostolika, […] & J. Burger, Early farmers from across Europe directly descended from Neolithic Aegeans, Proc. Natl. Acad. Sci. U.S.A.113 (25) 6886-6891, https://doi.org/10.1073/pnas.1523951113(2016).

[4] Hay, Maciamo, Haplogroups of Neolithic Europeans, Eupedia, https://www.eupedia.com/genetics/haplogroups_of_neolithic_farmers.shtml

Primorac D, Šarac J, Havaš Auguštin D, Novokmet N, Bego T, Pinhasi R, Šlaus M, Novak M, Marjanović D. Y Chromosome Story-Ancient Genetic Data as a Supplementary Tool for the Analysis of Modern Croatian Genetic Pool. Genes (Basel). 2024 Jun 6;15(6):748. doi: 10.3390/genes15060748. PMID: 38927684; PMCID: PMC11202852 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11202852/

António Faustino Carvalho, Eva Fernández-Domínguez, Eduardo Arroyo-Pardo, Catherine Robinson, João Luís Cardoso, João Zilhão, Mário Varela Gomes, Hunter-gatherer genetic persistence at the onset of megalithism in western Iberia: New mitochondrial evidence from Mesolithic and Neolithic necropolises in central-southern Portugal, Quaternary International, Volumes 677–678, 2023, Pages 111-120, ISSN 1040-6182, https://doi.org/10.1016/j.quaint.2023.03.015 .
(https://www.sciencedirect.com/science/article/pii/S104061822300099X )

[5] The Chalcolithic period, or Copper Age, is the phase that followed the Neolithic (New Stone Age) and preceded the Bronze Age. It is characterized by the discovery and use of copper metallurgy, marking a transition from the exclusive use of stone tools to the use of early metal tools alongside existing stone technology.This innovation led to advancements in agriculture, crafts, trade, and societal complexity, setting the stage for the more advanced Bronze Age societies

Chalcolithic, Wikipedia, This page was last edited on 19 September 2025, https://en.wikipedia.org/wiki/Chalcolithic

References for the facts discussed in the paragraph:

Rivollat, M.; Jeong, C.; Schiffels, S.; Kücükkalıpcı, İ.; Pemonge, M.-H.; Rohrlach, A. B.; Alt, K. W.; Binder, D.; Friederich, S.; Ghesquière, E.; Gronenborn, D.; Laporte, L.; Lefranc, P.; Meller, H.; Réveillas, H.; Rosenstock, E.; Rottier, S.; Scarre, C.; Soler, L.; Wahl, J.; Krause, J.; Deguilloux, M.-F.; Haak, W.: Ancient genome-wide DNA from France highlights the complexity of interactions between Mesolithic hunter-gatherers and Neolithic farmers. Science Advances 6 (22), eaaz5344 , pp. 1 – 16 (2020), https://www.science.org/doi/10.1126/sciadv.aaz5344

Arzelier, Ana & Rivollat, Maïté & Harmony, De & Marie-Hélène, Pemonge & Binder, Didier & Convertini, Fabien & Henri, Duday & Gandelin, Muriel & Jean, Guilaine & Wolfgang, Haak & Deguilloux, Marie-France & Pruvost, Melanie. (2022). Neolithic genomic data from Southern France showcase intensified interactions with hunter-gatherer communities. iScience. 25. 105387. 10.1016/j.isci.2022.105387 , https://www.researchgate.net/publication/364419697_Neolithic_genomic_data_from_Southern_France_showcase_intensified_interactions_with_hunter-gatherer_communities

Andrew Zeilstra and Johanna Knop, Heightened Interaction Between Neolithic Migrants and Hunter-Gatherers in Western Europe, 29 May 2020 Press Release, Max Planck Institte of Geoanthropology, https://www.shh.mpg.de/1713184/haak-french-dna#_ftnref4

Iñigo Olalde, et al. , Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups, bioRxiv 2025.03.24.644985; doi: https://doi.org/10.1101/2025.03.24.644985, https://www.biorxiv.org/content/10.1101/2025.03.24.644985v1.full; also found at: https://pubmed.ncbi.nlm.nih.gov/40196638

Tautalus, Geneticv Study: Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups, 25 Mar 2025, Forums > Population Genetics > Paleogenetics, Eupedia, https://www.eupedia.com/forum/threads/long-term-hunter-gatherer-continuity-in-the-rhine-meuse-region-was-disrupted-by-local-formation-of-expansive-bell-beaker-groups.45663/

Primorac D, Šarac J, Havaš Auguštin D, Novokmet N, Bego T, Pinhasi R, Šlaus M, Novak M, Marjanović D. Y Chromosome Story-Ancient Genetic Data as a Supplementary Tool for the Analysis of Modern Croatian Genetic Pool. Genes (Basel). 2024 Jun 6;15(6):748. doi: 10.3390/genes15060748. PMID: 38927684; PMCID: PMC11202852 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11202852/

Szécsényi-Nagy Anna, et al. Tracing the genetic origin of Europe’s first farmers reveals insights into their social organization. Proc Biol Sci. 2015 Apr 22;282(1805):20150339. doi: 10.1098/rspb.2015.0339. PMID: 25808890; PMCID: PMC4389623 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC4389623/

Wolfgang Haak, Oleg Balanovsky, Juan J. Sanchez, Sergey Koshel, Valery Zaporozhchenko, Christina J. Adler, Clio S. I. Der Sarkissian, Guido Brandt, Carolin Schwarz, Nicole Nicklisch, Veit Dresely, Barbara Fritsch, Elena Balanovska, Richard Villems, Harald Meller, Kurt W. Alt, Alan Cooper, Ancient DNA from European Early Neolithic Farmers Reveals Their Near Eastern Affinities, PLOS Biology, 9 Nov 2010, https://doi.org/10.1371/journal.pbio.1000536 

[6] Source of quote:

Hay, Maciamo, The great pairings of Y-DNA haplogroups in prehistory, 19 Jul 2015, Forums > Population Genetics > Y-DNA Haplogroups, Eupedia, https://www.eupedia.com/forum/threads/the-great-pairings-of-y-dna-haplogroups-in-prehistory.31431/

See also the following for explaining the transition to farming between hunter gatherer and early farming groups:

Leendert P. Louwe Kooijmans,” Transition to Farming Along the Lower Rhine and Meuse .” Ancient Europe, 8000 B.C. to A.D. 1000: Encyclopedia of the Barbarian World. . Encyclopedia.com. (September 3, 2025). https://www.encyclopedia.com/humanities/encyclopedias-almanacs-transcripts-and-maps/transition-farming-along-lower-rhine-and-meuse

[7] Kamjan S, Gillis RE, Çakırlar C, Raemaekers DCM. Specialized cattle farming in the Neolithic Rhine-Meuse Delta: Results from zooarchaeological and stable isotope (δ18O, δ13C, δ15N) analyses. PLoS One. 2020 Oct 21;15(10):e0240464. doi: 10.1371/journal.pone.0240464. PMID: 33085689; PMCID: PMC7577484. (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC7577484/

Olalde I, et al.  Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups. bioRxiv [Preprint]. 2025 Mar 25:2025.03.24.644985. doi: 10.1101/2025.03.24.644985. PMID: 40196638; PMCID: PMC11974744 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC11974744/

“Transition to Farming Along the Lower Rhine and Meuse .” Ancient Europe, 8000 B.C. to A.D. 1000: Encyclopedia of the Barbarian World. . Retrieved September 03, 2025 from Encyclopedia.com: https://www.encyclopedia.com/humanities/encyclopedias-almanacs-transcripts-and-maps/transition-farming-along-lower-rhine-and-meuse

Kamjan S, Gillis RE, Çakırlar C, Raemaekers DCM (2020) Specialized cattle farming in the Neolithic Rhine-Meuse Delta: Results from zooarchaeological and stable isotope (δ18O, δ13C, δ15N) analyses. PLoS ONE 15(10): e0240464. https://doi.org/10.1371/journal.pone.0240464 

[8] Iñigo Olalde, et al. , Long-term hunter-gatherer continuity in the Rhine-Meuse region was disrupted by local formation of expansive Bell Beaker groups, bioRxiv 2025.03.24.644985; doi:  https://doi.org/10.1101/2025.03.24.644985, https://www.biorxiv.org/content/10.1101/2025.03.24.644985v1.full; also found at: https://pubmed.ncbi.nlm.nih.gov/40196638

Kivisild T. The study of human Y chromosome variation through ancient DNA. Hum Genet. 2017 May;136(5):529-546. doi: 10.1007/s00439-017-1773-z. Epub 2017 Mar 4. Erratum in: Hum Genet. 2018 Oct;137(10):863. doi: 10.1007/s00439-018-1937-5. PMID: 28260210; PMCID: PMC5418327 (PubMed) https://pmc.ncbi.nlm.nih.gov/articles/PMC5418327/

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Click for Larger View | Source:Fokkens, Harry, The Periodisation of the Dutch Bronze Age: a critical review, in Metz, W.H. Beek, B.L. van Steegstra, H (ed.), Essays presented to Jay Jordan Butler on the occasion of his 80th birthday (pp.241-262) Metz, Van Beek & Steegstra , 2001,  uploaded to ResearchGate by Harry Fokkens on 14 October 2015, https://www.researchgate.net/publication/28646850_The_periodisation_of_the_Dutch_Bronze_Age_a_critical_review

Modified table from J.N. Lanting & W.G. Mook, The Pre- and Protohistory of the Netherlands in Terms of Radiocarbon Dates, Netherlands: Groningen, 1977

See also:

Fokkens, Harry, and Anthony Harding, ‘Introduction: The Bronze Age of Europe’, in Harry Fokkens, and Anthony Harding (eds), The Oxford Handbook of the European Bronze Age (2013; online edn, Oxford Academic, 5 Sept. 2013), https://doi.org/10.1093/oxfordhb/9780199572861.013.0001

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[38] A “star-like phylogeny” is a genetic tree showing a central node from which many short branches radiate, indicating a recent and rapid demographic expansion from a common ancestral lineage. This pattern is a classic indicator of a founder effect, a form of genetic bottleneck. 

Ailsa Allaby and Michael Allby “star phylogeny .” A Dictionary of Earth Sciences. . Encyclopedia.com. 2 Sep. 2025 . https://www.encyclopedia.com/earth-and-environment/ecology-and-environmentalism/environmental-studies/star-phylogeny#:~:text=star%20phylogeny%20In%20a%20phylogenetic,Modern%20Language%20Association

Founder effect

The founder effect occurs when a new population is established by a very small number of individuals from a larger population. This results in a new population with: 

  • Reduced genetic diversity: The founders carry only a fraction of the genetic variation present in the original population.
  • Unique genetic composition: The new population’s gene pool is a non-random sample of the source population’s genes. Some rare alleles may be over-represented, while others may be lost entirely. 

Ramachandran, S. , Deshpande, O. , Roseman, C. C. , Rosenberg, N. A. , Feldman, M. W. , & Cavalli‐Sforza, L. L. (2005). Support from the relationship of genetic and geographic distance in human populations for a serial founder effect originating in Africa. Proceedings of the National Academy of Sciences, 102, 15942–15947. 10.1073/pnas.0507611102 (PubMed) https://pubmed.ncbi.nlm.nih.gov/16243969/

How a star-like phylogeny is formed

  1. Founding event: A small group of individuals (the founders) with a limited gene pool moves to a new location.
  2. Population expansion: The new population grows rapidly, possibly due to a lack of competitors or predators.
  3. Low mutation accumulation: Since the population expanded very quickly, there has not been enough time for many new mutations to accumulate in the different lineages.
  4. Short, radiating branches: The minimal genetic differences among individuals result in a phylogeny with many short branches emanating from a single, central common ancestor (the founder or founder lineage). 

The star-like pattern provides two key pieces of evidence for recent and rapid growth: 

  • Short branches: The short length of the branches indicates that very few mutations occurred in each lineage since the founding event, suggesting that not much time has passed. 
  • Common ancestor: All individuals can trace their ancestry back to a single, recent founder lineage, represented by the central node.